EcoService Models Library (ESML)
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Compare EMs
Which comparison is best for me?EM Variables by Variable Role
One quick way to compare ecological models (EMs) is by comparing their variables. Predictor variables show what kinds of influences a model is able to account for, and what kinds of data it requires. Response variables show what information a model is capable of estimating.
This first comparison shows the names (and units) of each EM’s variables, side-by-side, sorted by variable role. Variable roles in ESML are as follows:
- Predictor Variables
- Time- or Space-Varying Variables
- Constants and Parameters
- Intermediate (Computed) Variables
- Response Variables
- Computed Response Variables
- Measured Response Variables
EM Variables by Category
A second way to use variables to compare EMs is by focusing on the kind of information each variable represents. The top-level categories in the ESML Variable Classification Hierarchy are as follows:
- Policy Regarding Use or Management of Ecosystem Resources
- Land Surface (or Water Body Bed) Cover, Use or Substrate
- Human Demographic Data
- Human-Produced Stressor or Enhancer of Ecosystem Goods and Services Production
- Ecosystem Attributes and Potential Supply of Ecosystem Goods and Services
- Non-monetary Indicators of Human Demand, Use or Benefit of Ecosystem Goods and Services
- Monetary Values
Besides understanding model similarities, sorting the variables for each EM by these 7 categories makes it easier to see if the compared models can be linked using similar variables. For example, if one model estimates an ecosystem attribute (in Category 5), such as water clarity, as a response variable, and a second model uses a similar attribute (also in Category 5) as a predictor of recreational use, the two models can potentially be used in tandem. This comparison makes it easier to spot potential model linkages.
All EM Descriptors
This selection allows a more detailed comparison of EMs by model characteristics other than their variables. The 50-or-so EM descriptors for each model are presented, side-by-side, in the following categories:
- EM Identity and Description
- EM Modeling Approach
- EM Locations, Environments, Ecology
- EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
EM Descriptors by Modeling Concepts
This feature guides the user through the use of the following seven concepts for comparing and selecting EMs:
- Conceptual Model
- Modeling Objective
- Modeling Context
- Potential for Model Linkage
- Feasibility of Model Use
- Model Certainty
- Model Structural Information
Though presented separately, these concepts are interdependent, and information presented under one concept may have relevance to other concepts as well.
EM Identity and Description
EM ID
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EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
EM Short Name
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Green biomass production, Central French Alps | Community flowering date, Central French Alps | C Sequestration and De-N, Tampa Bay, FL, USA | SAV occurrence, St. Louis River, MN/WI, USA | C sequestration in grassland restoration, England | WESP: Irrigation water, ID, USA |
EM Full Name
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Green biomass production, Central French Alps | Community weighted mean flowering date, Central French Alps | Value of Carbon Sequestration and Denitrification benefits, Tampa Bay, FL, USA | Predicting submerged aquatic vegetation occurrence, St. Louis River Estuary, MN & WI, USA | Carbon sequestration in grassland diversity restoration, England | WESP: Irrigation return water treatment, Idaho, USA |
EM Source or Collection
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EU Biodiversity Action 5 | EU Biodiversity Action 5 | US EPA | US EPA | None | None |
EM Source Document ID
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260 | 260 | 186 | 330 | 396 |
393 ?Comment:Additional data came from electronic appendix provided by author Chris Murphy. |
Document Author
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Lavorel, S., Grigulis, K., Lamarque, P., Colace, M-P, Garden, D., Girel, J., Pellet, G., and Douzet, R. | Lavorel, S., Grigulis, K., Lamarque, P., Colace, M-P, Garden, D., Girel, J., Pellet, G., and Douzet, R. | Russell, M. and Greening, H. | Ted R. Angradi, Mark S. Pearson, David W. Bolgrien, Brent J. Bellinger, Matthew A. Starry, Carol Reschke | De Deyn, G. B., R. S. Shiel, N. J. Ostle, N. P. McNamara, S. Oakley, I. Young, C. Freeman, N. Fenner, H. Quirk, and R. D. Bardgett | Murphy, C. and T. Weekley |
Document Year
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2011 | 2011 | 2013 | 2013 | 2011 | 2012 |
Document Title
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Using plant functional traits to understand the landscape distribution of multiple ecosystem services | Using plant functional traits to understand the landscape distribution of multiple ecosystem services | Estimating benefits in a recovering estuary: Tampa Bay, Florida | Predicting submerged aquatic vegetation cover and occurrence in a Lake Superior estuary | Additional carbon sequestration benefits of grassland diversity restoration | Measuring outcomes of wetland restoration, enhancement, and creation in Idaho-- Assessing potential functions, values, and condition in a watershed context. |
Document Status
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Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published |
Comments on Status
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Published journal manuscript | Published journal manuscript | Published journal manuscript | Published journal manuscript | Published journal manuscript | Published report |
EM ID
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EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | |
Contact Name
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Sandra Lavorel | Sandra Lavorel | M. Russell | Ted R. Angradi | Gerlinde B. De Deyn | Chris Murphy |
Contact Address
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Laboratoire d’Ecologie Alpine, UMR 5553 CNRS Université Joseph Fourier, BP 53, 38041 Grenoble Cedex 9, France | Laboratoire d’Ecologie Alpine, UMR 5553 CNRS Université Joseph Fourier, BP 53, 38041 Grenoble Cedex 9, France | US EPA, Gulf Ecology Division, 1 Sabine Island Dr, Gulf Breeze, FL 32563, USA | U.S. Environmental Protection Agency, Office of Research and Development, National Health and Environmental Effects Research Laboratory, Mid-Continent Ecology Division, 6201 Congdon Blvd., Duluth, MN 55804, USA | Dept. of Terrestrial Ecology, Netherlands Institute of Ecology, P O Box 40, 6666 ZG Heteren, The Netherlands | Idaho Dept. Fish and Game, Wildlife Bureau, Habitat Section, Boise, ID |
Contact Email
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sandra.lavorel@ujf-grenoble.fr | sandra.lavorel@ujf-grenoble.fr | Russell.Marc@epamail.epa.gov | angradi.theodore@epa.gov | g.dedeyn@nioo.knaw.nl; gerlindede@gmail.com | chris.murphy@idfg.idaho.gov |
EM ID
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EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
Summary Description
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ABSTRACT: "Here, we propose a new approach for the analysis, mapping and understanding of multiple ES delivery in landscapes. Spatially explicit single ES models based on plant traits and abiotic characteristics are combined to identify ‘hot’ and ‘cold’ spots of multiple ES delivery, and the land use and biotic determinants of such distributions. We demonstrate the value of this trait-based approach as compared to a pure land-use approach for a pastoral landscape from the central French Alps, and highlight how it improves understanding of ecological constraints to, and opportunities for, the delivery of multiple services. Vegetative height and leaf traits such as leaf dry matter content were response traits strongly influenced by land use and abiotic environment, with follow-on effects on several ecosystem properties (e.g., green biomass production), and could therefore be used as functional markers of ES." AUTHOR'S DESCRIPTION: "Variation in green biomass production was modelled using…traits community-weighted mean (CWM) and functional divergence (FD) and abiotic variables (continuous variables; trait + abiotic) following Diaz et al. (2007). …The comparison between this model and the land-use alone model identifies the need for site-based information beyond a land use or land cover proxy, and the comparison with the land use + abiotic model assesses the value of additional ecological (trait) information…Green biomass production for each pixel was calculated and mapped using model estimates for…regression coefficients on abiotic variables and traits. For each pixel these calculations were applied to mapped estimates of abiotic variables and trait CWM and FD. This step is critically novel as compared to a direct application of the model by Diaz et al. (2007) in that we explicitly modelled the responses of trait community-weighted means and functional divergences to environment prior to evaluating their effects on ecosystem properties. Such an approach is the key to the explicit representation of functional variation across the landscape, as opposed to the use of unique trait values within each land use (see Albert et al. 2010)." | ABSTRACT: "Here, we propose a new approach for the analysis, mapping and understanding of multiple ES delivery in landscapes. Spatially explicit single ES models based on plant traits and abiotic characteristics are combined to identify ‘hot’ and ‘cold’ spots of multiple ES delivery, and the land use and biotic determinants of such distributions. We demonstrate the value of this trait-based approach as compared to a pure land-use approach for a pastoral landscape from the central French Alps, and highlight how it improves understanding of ecological constraints to, and opportunities for, the delivery of multiple services." AUTHOR'S DESCRIPTION: "Community-weighted mean date of flowering onset was modelled using mixed models with land use and abiotic variables as fixed effects (LU + abiotic model) and year as a random effect…and modelled for each 20 x 20 m pixel using GLM estimated effects for each land use category and estimated regression coefficients with abiotic variables." | AUTHOR'S DESCRIPTION: "...we examine the change in the production of ecosystem goods produced as a result of restoration efforts and potential relative cost savings for the Tampa Bay community from seagrass expansion (more than 3,100 ha) and coastal marsh and mangrove restoration (∼600 ha), since 1990… The objectives of this article are to explore the roles that ecological processes and resulting ecosystem goods have in maintaining healthy estuarine systems by (1) quantifying the production of specific ecosystem goods in a subtropical estuarine system and (2) determining potential cost savings of improved water quality and increased habitat in a recovering estuary." (pp. 2) | ABSTRACT: “Submerged aquatic vegetation (SAV) provides the biophysical basis for multiple ecosystem services in Great Lakes estuaries. Understanding sources of variation in SAV is necessary for sustainable management of SAV habitat. From data collected using hydroacoustic survey methods, we created predictive models for SAV in the St. Louis River Estuary (SLRE) of western Lake Superior. The dominant SAV species in most areas of the estuary was American wild celery (Vallisneria americana Michx.)…” AUTHOR’S DESCRIPTION: “The SLRE is a Great Lakes “rivermouth” ecosystem as defined by Larson et al. (2013). The 5000-ha estuary forms a section of the state border between Duluth, Minnesota and Superior, Wisconsin…In the SLRE, SAV beds are often patchy, turbidity varies considerably among areas (DeVore, 1978) and over time, and the growing season is short. Given these conditions, hydroacoustic survey methods were the best option for generating the extensive, high resolution data needed for modeling. From late July through mid September in 2011, we surveyed SAV in Allouez Bay, part of Superior Bay, eastern half of St. Louis Bay, and Spirit Lake…We used the measured SAV percent cover at the location immediately previous to each useable record location along each transect as a lag variable to correct for possible serial autocorrelation of model error. SAV percent cover, substrate parameters, corrected depth, and exposure and bed slope data were combined in Arc-GIS...We created logistic regression models for each area of the SLRE to predict the probability of SAV being present at each report location. We created models for the training data set using the Logistic procedure in SAS v.9.1 with step wise elimination (?=0.05). Plots of cover by depth for selected predictor values (Supplementary Information Appendix C) suggested that interactions between depth and other predictors were likely to be significant, and so were included in regression models. We retained the main effect if their interaction terms were significant in the model. We examined the performance of the models using the area under the receiver operating characteristic (AUROC) curve. AUROC is the probability of concordance between random pairs of observations and ranges from 0.5 to 1 (Gönen, 2006). We cross-validated logistic occurrence models for their ability to classify correctly locations in the validation (holdout) dataset and in the Superior Bay dataset… Model performance, as indicated by the area under the receiver operating characteristic (AUROC) curve was >0.8 (Table 3). Assessed accuracy of models (the percent of records where the predicted probability of occurrence and actual SAV presence or absence agreed) for split datasets was 79% for Allouez Bay, 86% for St. Louis Bay, and 78% for Spirit Lake." | ABSTRACT: "A major aim of European agri-environment policy is the management of grassland for botanical diversity conservation and restoration, together with the delivery of ecosystem services including soil carbon (C) sequestration. To test whether management for biodiversity restoration has additional benefits for soil C sequestration, we investigated C and nitrogen (N) accumulation rates in soil and C and N pools in vegetation in a long-term field experiment (16 years) in which fertilizer application and plant seeding were manipulated. In addition, the abundance of the legume Trifolium pratense was manipulated for the last 2 years. To unravel the mechanisms underlying changes in soil C and N pools, we also tested for effects of diversity restoration management on soil structure, ecosystem respiration and soil enzyme activities…" AUTHOR'S DESCRIPTION: "Measurements were made on 36 plots of 3 x 3 m comprising two management treatments (and their controls) in a long-term multifactorial grassland restoration experiment which have successfully increased plant species diversity, namely the cessation of NPK fertilizer application and the addition of seed mixtures…" | A wetland restoration monitoring and assessment program framework was developed for Idaho. The project goal was to assess outcomes of substantial governmental and private investment in wetland restoration, enhancement and creation. The functions, values, condition, and vegetation at restored, enhanced, and created wetlands on private and state lands across Idaho were retrospectively evaluated. Assessment was conducted at multiple spatial scales and intensities. Potential functions and values (ecosystem services) were rapidly assessed using the Oregon Rapid Wetland Assessment Protocol. Vegetation samples were analyzed using Floristic Quality Assessment indices from Washington State. We compared vegetation of restored, enhanced, and created wetlands with reference wetlands that occurred in similar hydrogeomorphic environments determined at the HUC 12 level. |
Specific Policy or Decision Context Cited
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None identified | None identified | Restoration of seagrass | None identified | None identified | None identified |
Biophysical Context
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Elevation ranges from 1552 to 2442 m, on predominately south-facing slopes | Elevation ranges from 1552 to 2442 m, on predominantly south-facing slopes | Recovering estuary; Seagrass; Coastal fringe; Saltwater marsh; Mangrove | submerged aquatic vegetation | Lolium perenne-Cynosorus cristatus grassland; The soil is a shallow brown-earth (average depth 28 cm) over limestone of moderate-high residual fertility. | restored, enhanced and created wetlands |
EM Scenario Drivers
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No scenarios presented | No scenarios presented | Habitat loss or restoration in Tampa Bay Estuary | No scenarios presented | Additional benefits due to biodiversity restoration practices | Sites, function or habitat focus |
EM ID
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EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
Method Only, Application of Method or Model Run
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Method + Application | Method + Application | Method + Application | Method + Application | Method + Application (multiple runs exist) View EM Runs | Method + Application (multiple runs exist) View EM Runs |
New or Pre-existing EM?
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New or revised model | New or revised model | New or revised model | New or revised model | New or revised model | New or revised model |
Related EMs (for example, other versions or derivations of this EM) described in ESML
EM ID
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EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
Document ID for related EM
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Doc-260 | Doc-260 | Doc-269 | None | None | None | Doc-390 |
EM ID for related EM
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EM-66 | EM-68 | EM-69 | EM-70 | EM-71 | EM-79 | EM-80 | EM-81 | EM-82 | EM-83 | EM-65 | EM-66 | EM-68 | EM-69 | EM-70 | EM-79 | EM-80 | EM-81 | EM-82 | EM-83 | None | None | None | EM-718 | EM-734 | EM-760 | EM-761 | EM-763 | EM-764 | EM-766 | EM-767 | EM-768 |
EM Modeling Approach
EM ID
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EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
EM Temporal Extent
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2007-2009 | 2007-2008 | 1982-2010 | 2010 - 2012 | 1990-2007 | 2010-2012 |
EM Time Dependence
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time-stationary | time-stationary | time-stationary | time-stationary | time-stationary | time-dependent |
EM Time Reference (Future/Past)
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Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | past time |
EM Time Continuity
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Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable |
EM Temporal Grain Size Value
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Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable |
EM Temporal Grain Size Unit
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Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable |
EM ID
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EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
Bounding Type
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Physiographic or Ecological | Physiographic or Ecological | Physiographic or Ecological | Physiographic or ecological | Other | Multiple unrelated locations (e.g., meta-analysis) |
Spatial Extent Name
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Central French Alps | Central French Alps | Tampa Bay Estuary | St. Louis River Estuary | Colt Park meadows, Ingleborough National Nature Reserve, northern England | Wetlands in idaho |
Spatial Extent Area (Magnitude)
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10-100 km^2 | 10-100 km^2 | 1000-10,000 km^2. | 10-100 km^2 | <1 ha | 100,000-1,000,000 km^2 |
EM ID
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EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
EM Spatial Distribution
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spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially distributed (in at least some cases) |
spatially distributed (in at least some cases) ?Comment:BH: Each individual transect?s data was parceled into location reports, and that each report?s ?quadrat? area was dependent upon the angle of the hydroacoustic sampling beam. The spatial grain is 0.07 m^2, 0.20 m^2 and 0.70 m^2 for depths of 1 meter, 2 meters and 3 meters, respectively. |
spatially distributed (in at least some cases) | spatially lumped (in all cases) |
Spatial Grain Type
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area, for pixel or radial feature | area, for pixel or radial feature | area, for pixel or radial feature | area, for pixel or radial feature | area, for pixel or radial feature | Not applicable |
Spatial Grain Size
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20 m x 20 m | 20 m x 20 m | 1 ha | 0.07 m^2 to 0.70 m^2 | 3 m x 3 m | Not applicable |
EM ID
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EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
EM Computational Approach
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Analytic | Analytic | Analytic | Analytic | Analytic | Numeric |
EM Determinism
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deterministic | deterministic | deterministic | deterministic | stochastic | deterministic |
Statistical Estimation of EM
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EM ID
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EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
Model Calibration Reported?
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No | No | Yes | Yes | Not applicable | No |
Model Goodness of Fit Reported?
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Yes | Yes | No | Yes | Not applicable | No |
Goodness of Fit (metric| value | unit)
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None |
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None | None |
Model Operational Validation Reported?
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Yes | No | No | Yes | No | No |
Model Uncertainty Analysis Reported?
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No | No | No | No | No | No |
Model Sensitivity Analysis Reported?
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No | No | No | No | No | No |
Model Sensitivity Analysis Include Interactions?
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Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable |
EM Locations, Environments, Ecology
Terrestrial location (Classification hierarchy: Continent > Country > U.S. State [United States only])
EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
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None |
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Marine location (Classification hierarchy: Realm > Region > Province > Ecoregion)
EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
None | None |
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None | None | None |
Centroid Lat/Long (Decimal Degree)
EM ID
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EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
Centroid Latitude
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45.05 | 45.05 | 27.95 | 46.72 | 54.2 | 44.06 |
Centroid Longitude
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6.4 | 6.4 | -82.47 | -96.13 | -2.35 | -114.69 |
Centroid Datum
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WGS84 | WGS84 | WGS84 | WGS84 | WGS84 | WGS84 |
Centroid Coordinates Status
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Provided | Provided | Estimated | Estimated | Provided | Estimated |
EM ID
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EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
EM Environmental Sub-Class
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Agroecosystems | Grasslands | Agroecosystems | Grasslands | Near Coastal Marine and Estuarine | Aquatic Environment (sub-classes not fully specified) | Rivers and Streams | Inland Wetlands | Lakes and Ponds | Agroecosystems | Grasslands | Inland Wetlands |
Specific Environment Type
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Subalpine terraces, grasslands, and meadows | Subalpine terraces, grasslands, and meadows. | Subtropical Estuary | Freshwater estuarine system | fertilized grassland (historically hayed) | created, restored and enhanced wetlands |
EM Ecological Scale
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Not applicable | Not applicable | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class |
Scale of differentiation of organisms modeled
EM ID
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EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
EM Organismal Scale
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Community | Community | Not applicable | Not applicable | Community | Not applicable |
Taxonomic level and name of organisms or groups identified
EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
None Available | None Available | None Available | None Available | None Available | None Available |
EnviroAtlas URL
EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
CICES v 4.3 - Common International Classification of Ecosystem Services (Section > Division > Group > Class)
EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
None | None |
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<a target="_blank" rel="noopener noreferrer" href="https://www.epa.gov/eco-research/national-ecosystem-services-classification-system-nescs-plus">National Ecosystem Services Classification System (NESCS) Plus</a>
(Environmental Subclass > Ecological End-Product (EEP) > EEP Subclass > EEP Modifier)
EM-65 | EM-71 | EM-195 | EM-414 |
EM-735 ![]() |
EM-743 ![]() |
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None |
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None | None | None |