{PAGE NAME} | {WEB AREA NAME}

EM Identity and Description

EM Identification (* Note that run information is shown only where run data differ from the "parent" entry shown at left)

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
EM Short Name em.detail.shortNameHelp ?	Fish nutrient cycling , Ohio, USA	*	*	*	*	*	*	*
EM Full Name em.detail.fullNameHelp ?	Nutrient Cycling by gizzard shad, Ohio, USA	*	*	*	*	*	*	*
EM Source or Collection em.detail.emSourceOrCollectionHelp ?	None	*	*	*	*	*	*	*
EM Source Document ID	385	*	*	*	*	*	*	*
Document Author em.detail.documentAuthorHelp ?	Vanni, M.J., Bowling, A.M., Dickman, E.M., Hale, R.S., Higgins, K.A., Horgan, M.J., Knoll, L.B., Renwick, W.H., and R.A. Stein	*	*	*	*	*	*	*
Document Year em.detail.documentYearHelp ?	2006	*	*	*	*	*	*	*
Document Title em.detail.sourceIdHelp ?	Nutrient cycling by fish supports relatively more primary production as lake productivity increases	*	*	*	*	*	*	*
Document Status em.detail.statusCategoryHelp ?	Peer reviewed and published	*	*	*	*	*	*	*
Comments on Status em.detail.commentsOnStatusHelp ?	Published journal manuscript	*	*	*	*	*	*	*

Software and Access (* Note that run information is shown only where run data differ from the "parent" entry shown at left)

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
EM Software Access info Exit em.detail.modelAccessInformationHelp ?	Not applicable	*	*	*	*	*	*	*
Contact Name em.detail.contactNameHelp ?	Michael Vanni	*	*	*	*	*	*	*
Contact Address	Dept of Environmental toxocology, C.emson Univ. Pendleton, South Carolina 29670, USA	*	*	*	*	*	*	*
Contact Email	vannimj@muohio.edu	*	*	*	*	*	*	*

EM Description (* Note that run information is shown only where run data differ from the "parent" entry shown at left)

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
Summary Description em.detail.summaryDescriptionHelp ?	ABSTRACT: "Animals can be important in nutrient cycling in particular ecosystems, but few studies have examined how this importance varies along environmental gradients. In this study we quantified the nutrient cycling role of an abundant detritivorous fish species, the gizzard shad (Dorosoma cepedianum), in reservoir ecosystems along a gradient of ecosystem productivity. Gizzard shad feed mostly on sediment detritus and excrete sediment-derived nutrients into the water column, thereby mediating a cross-habitat translocation of nutrients to phytoplankton. We quantified nitrogen and phosphorus cycling (excretion) rates of gizzard shad, as well as nutrient demand by phytoplankton, in seven lakes over a four-year period (16 lake-years). The lakes span a gradient of watershed land use (the relative amounts of land used for agriculture vs. forest) and productivity. As the watersheds of these lakes became increasingly dominated by agricultural land, primary production rates, lake trophic state indicators (total phosphorus and chlorophyll concentrations), and nutrient flux through gizzard shad populations all increased. Nutrient cycling by gizzard shad supported a substantial proportion of primary production in these ecosystems, and this proportion increased as watershed agriculture (and ecosystem productivity) increased. In the four productive lakes with agricultural watersheds (.78% agricultural land), gizzard shad supported on average 51% of phytoplankton primary production (range 27–67%). In contrast, in the three relatively unproductive lakes in forested or mixed-land-use watersheds (.47% forest, ,52% agricultural land), gizzard shad supported 18% of primary production (range 14–23%). Thus, along a gradient of forested to agricultural landscapes, both watershed nutrient inputs and nutrient translocation by gizzard shad increase, but our data indicate that the importance of nutrient translocation by gizzard shad increases more rapidly. Our results therefore support the hypothesis that watersheds and gizzard shad jointly regulate primary production in reservoir ecosystems "	*	*	*	*	*	*	*
Specific Policy or Decision Context Cited em.detail.policyDecisionContextHelp ?	None identified	*	*	*	*	*	*	*
Biophysical Context	Lakes	*	*	*	*	*	*	*
EM Scenario Drivers em.detail.scenarioDriverHelp ?	Lake productivity	*	*	*	*	*	*	*

EM Relationship to Other EMs or Applications

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
Method Only, Application of Method or Model Run em.detail.methodOrAppHelp ?	Method + Application (multiple runs exist)	Model Run Associated with a Specific EM Application	Model Run Associated with a Specific EM Application	Model Run Associated with a Specific EM Application	Model Run Associated with a Specific EM Application	Model Run Associated with a Specific EM Application	Model Run Associated with a Specific EM Application	Model Run Associated with a Specific EM Application
New or Pre-existing EM? em.detail.newOrExistHelp ?	New or revised model	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake

Related EMs (for example, other versions or derivations of this EM) described in ESML

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
Document ID for related EM em.detail.relatedEmDocumentIdHelp ?	None	None	None	None	None	None	None	None
EM ID for related EM em.detail.relatedEmEmIdHelp ?	None	None	None	None	None	None	None	None

EM Modeling Approach

EM Relationship to Time (* Note that run information is shown only where run data differ from the "parent" entry shown at left)

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
EM Temporal Extent em.detail.tempExtentHelp ?	2000-2003	*	*	*	*	*	*	*
EM Time Dependence em.detail.timeDependencyHelp ?	time-stationary	*	*	*	*	*	*	*
EM Time Reference (Future/Past) em.detail.futurePastHelp ?	Not applicable	*	*	*	*	*	*	*
EM Time Continuity em.detail.continueDiscreteHelp ?	Not applicable	*	*	*	*	*	*	*
EM Temporal Grain Size Value em.detail.tempGrainSizeHelp ?	Not applicable	*	*	*	*	*	*	*
EM Temporal Grain Size Unit em.detail.tempGrainSizeUnitHelp ?	Not applicable	*	*	*	*	*	*	*

EM spatial extent (* Note that run information is shown only where run data differ from the "parent" entry shown at left)

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
Bounding Type em.detail.boundingTypeHelp ?	Multiple unrelated locations (e.g., meta-analysis)	*	*	*	*	*	*	*
Spatial Extent Name em.detail.extentNameHelp ?	Lakes in Ohio	*	*	*	*	*	*	*
Spatial Extent Area (Magnitude) em.detail.extentAreaHelp ?	100,000-1,000,000 km^2	*	*	*	*	*	*	*

Spatial Distribution of Computations (* Note that run information is shown only where run data differ from the "parent" entry shown at left)

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
EM Spatial Distribution em.detail.distributeLumpHelp ?	spatially lumped (in all cases)	*	*	*	*	*	*	*
Spatial Grain Type em.detail.spGrainTypeHelp ?	Not applicable	*	*	*	*	*	*	*
Spatial Grain Size em.detail.spGrainSizeHelp ?	Not applicable	*	*	*	*	*	*	*

EM Structure and Computation Approach (* Note that run information is shown only where run data differ from the "parent" entry shown at left)

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
EM Computational Approach em.detail.emComputationalApproachHelp ?	Numeric	*	*	*	*	*	*	*
EM Determinism em.detail.deterStochHelp ?	deterministic	*	*	*	*	*	*	*
Statistical Estimation of EM em.detail.statisticalEstimationHelp ?	Conventional (frequentist) statistics	*	*	*	*	*	*	*

Model Checking Procedures Used (* Note that run information is shown only where run data differ from the "parent" entry shown at left)

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
Model Calibration Reported? em.detail.calibrationHelp ?	Yes ? Comment: Nitrogen and Phosphorus excretion rates were calibrated by lake and fish size class.	* ? Comment: Nitrogen and Phosphorus excretion rates were calibrated by lake and fish size class.	* ? Comment: Nitrogen and Phosphorus excretion rates were calibrated by lake and fish size class.	* ? Comment: Nitrogen and Phosphorus excretion rates were calibrated by lake and fish size class.	* ? Comment: Nitrogen and Phosphorus excretion rates were calibrated by lake and fish size class.	* ? Comment: Nitrogen and Phosphorus excretion rates were calibrated by lake and fish size class.	* ? Comment: Nitrogen and Phosphorus excretion rates were calibrated by lake and fish size class.	* ? Comment: Nitrogen and Phosphorus excretion rates were calibrated by lake and fish size class.
Model Goodness of Fit Reported? em.detail.goodnessFitHelp ?	No	*	*	*	*	*	*	*
Goodness of Fit (metric\| value \| unit) em.detail.goodnessFitValuesHelp ?	None	*	*	*	*	*	*	*
Model Operational Validation Reported? em.detail.validationHelp ?	No	*	*	*	*	*	*	*
Model Uncertainty Analysis Reported? em.detail.uncertaintyAnalysisHelp ?	No	*	*	*	*	*	*	*
Model Sensitivity Analysis Reported? em.detail.sensAnalysisHelp ?	No	*	*	*	*	*	*	*
Model Sensitivity Analysis Include Interactions? em.detail.interactionConsiderHelp ?	Not applicable	*	*	*	*	*	*	*

EM Locations, Environments, Ecology

Location of EM Application (* Note that run information is shown only where run data differ from the "parent" entry shown at left)

Terrestrial location (Classification hierarchy: Continent > Country > U.S. State [United States only])

EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
North America United States Ohio	*	*	*	*	*	*	*

Marine location (Classification hierarchy: Realm > Region > Province > Ecoregion)

EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
None	*	*	*	*	*	*	*

Centroid Lat/Long (Decimal Degree)

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
Centroid Latitude em.detail.ddLatHelp ?	40.15	*	*	*	*	*	*	*
Centroid Longitude em.detail.ddLongHelp ?	-82.95	*	*	*	*	*	*	*
Centroid Datum em.detail.datumHelp ?	WGS84	*	*	*	*	*	*	*
Centroid Coordinates Status em.detail.coordinateStatusHelp ?	Estimated	*	*	*	*	*	*	*

Environments and Scales Modeled (* Note that run information is shown only where run data differ from the "parent" entry shown at left)

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
EM Environmental Sub-Class em.detail.emEnvironmentalSubclassHelp ?	Lakes and Ponds	*	*	*	*	*	*	*
Specific Environment Type em.detail.specificEnvTypeHelp ?	Reservoirs	*	*	*	*	*	*	*
EM Ecological Scale em.detail.ecoScaleHelp ?	Ecological scale corresponds to the Environmental Sub-class	*	*	*	*	*	*	*

Organisms modeled (* Note that run information is shown only where run data differ from the "parent" entry shown at left)

Scale of differentiation of organisms modeled

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
EM Organismal Scale em.detail.orgScaleHelp ?	Not applicable	*	*	*	*	*	*	*

Taxonomic level and name of organisms or groups identified

EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
Species Dorosoma cepedianum (Gizzard Shad)	*	*	*	*	*	*	*

EnviroAtlas URL

EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
Average Annual Precipitation, Waterbody area, The Watershed Boundary Dataset (WBD)	Average Annual Precipitation, Waterbody area, The Watershed Boundary Dataset (WBD)	Average Annual Precipitation, Waterbody area, The Watershed Boundary Dataset (WBD)	Average Annual Precipitation, Waterbody area, The Watershed Boundary Dataset (WBD)	Average Annual Precipitation, Waterbody area, The Watershed Boundary Dataset (WBD)	Average Annual Precipitation, Waterbody area, The Watershed Boundary Dataset (WBD)	Average Annual Precipitation, Waterbody area, The Watershed Boundary Dataset (WBD)	Average Annual Precipitation, Waterbody area, The Watershed Boundary Dataset (WBD)

EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system

* Note that run information is shown only where run data differ from the "parent" entry shown at left

CICES v 4.3 - Common International Classification of Ecosystem Services (Section > Division > Group > Class)

EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
None	*	*	*	*	*	*	*

National Ecosystem Services Classification System (NESCS) Plus

(Environmental Subclass > Ecological End-Product (EEP) > EEP Subclass > EEP Modifier)

EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
None	*	*	*	*	*	*	*

EM Variable Names (and Units)

* Note that for runs, variable name is displayed only where data for that variable differed by run AND those differences were reported in the source document. Where differences occurred but were not reported, the variable is not listed. Click on variable name to view details.

To view run information for variables, select any variable below or view run data for all variables.

Predictor

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
Driving Variables (and Units) em.detail.drivingVariableHelp ?	Chlorophyll a concentration (ug/L) Fish lengths (mm) Lake class (Not applicable) Land use (Not applicable) Nitrogen excretion rate (per capita/size class) for gizzard shad (ug N/fish/day) Phosphorus excretion rate (per capitat/size class) for gizzard shad (ug P/fish/day) Proportion of total fish that are gizzard shad (%) Seston carbon concentration (umol/L) Seston nitrogen concentration (umol/L) Seston phosphorus concentration (μmol/L) Solar irradiance (W/m^2) Total fish (No./m^2) Total phosphorus (ug P/L)	Chlorophyll a concentration (ug/L) Phosphorus excretion rate (per capitat/size class) for gizzard shad (μg P/fish/day) Seston carbon concentration (umol/L) Seston nitrogen concentration (umol/L) Seston phosphorus concentration (μmol/L) Total phosphorus (ug P/L)	Chlorophyll a concentration (ug/L) Lake class (Piedmont lake) (Not applicable) Phosphorus excretion rate (per capitat/size class) for gizzard shad (μg P/fish/day) Seston carbon concentration (umol/L) Seston nitrogen concentration (umol/L) Seston phosphorus concentration (μmol/L) Total phosphorus (ug P/L)	Chlorophyll a concentration (ug/L) Lake class (Pleasant Hill) (Not applicable) Phosphorus excretion rate (per capitat/size class) for gizzard shad (μg P/fish/day) Seston carbon concentration (umol/L) Seston nitrogen concentration (umol/L) Seston phosphorus concentration (μmol/L) Total phosphorus (ug P/L)	Chlorophyll a concentration (ug/L) Lake class (Kiser) (Not applicable) Phosphorus excretion rate (per capitat/size class) for gizzard shad (μg P/fish/day) Seston carbon concentration (umol/L) Seston nitrogen concentration (umol/L) Seston phosphorus concentration (μmol/L) Total phosphorus (ug P/L)	Chlorophyll a concentration (ug/L) Lake class (O'Shaughnessy) (Not applicable) Phosphorus excretion rate (per capitat/size class) for gizzard shad (μg P/fish/day) Seston carbon concentration (umol/L) Seston nitrogen concentration (umol/L) Seston phosphorus concentration (μmol/L) Total phosphorus (ug P/L)	Chlorophyll a concentration (ug/L) Lake class (C.J. Brown) (Not applicable) Phosphorus excretion rate (per capitat/size class) for gizzard shad (μg P/fish/day) Seston carbon concentration (umol/L) Seston nitrogen concentration (umol/L) Seston phosphorus concentration (μmol/L) Total phosphorus (ug P/L)	Chlorophyll a concentration (ug/L) Lake class (Acton) (Not applicable) Phosphorus excretion rate (per capitat/size class) for gizzard shad (μg P/fish/day) Seston carbon concentration (umol/L) Seston nitrogen concentration (umol/L) Seston phosphorus concentration (μmol/L) Total phosphorus (ug P/L)
Constant or Factor Variables (and Units) em.detail.constantFactorVariableHelp ?	Size class (gizzard shad) (Not applicable) Watershed boundary (Not applicable)	Lake class (Burr Oak) (Not applicable)	*	*	*	*	*	*

Intermediate

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
Intermediate (Computed) Variables (and Units) em.detail.intermediateVariableHelp ?	Excretion rate per size class (μg N/g fish) P excretion rate (per size class) (μg P/g fish) Photosynthetically active radiation (PAR) (Not reported) Primary production rate (mg C/m^2/day) Seston C:N (molar) Seston C:P (molar)	N excretion rate per size class (μg N/g fish) P excretion rate (per size class) (μg P/g fish) Primary production rate (mg C/m^2/day) Seston C:N (molar) Seston C:P (molar)	N excretion rate per size class (μg N/g fish) P excretion rate (per size class) (μg P/g fish) Primary production rate (mg C/m^2/day) Seston C:N (molar) Seston C:P (molar)	N excretion rate per size class (μg N/g fish) P excretion rate (per size class) (μg P/g fish) Primary production rate (mg C/m^2/day) Seston C:N (molar) Seston C:P (molar)	N excretion rate per size class (μg N/g fish) P excretion rate (per size class) (μg P/g fish) Primary production rate (mg C/m^2/day) Seston C:N (molar) Seston C:P (molar)	N excretion rate per size class (μg N/g fish) P excretion rate (per size class) (μg P/g fish) Primary production rate (mg C/m^2/day) Seston C:N (molar) Seston C:P (molar)	N excretion rate per size class (μg N/g fish) P excretion rate (per size class) (μg P/g fish) Primary production rate (mg C/m^2/day) Seston C:N (molar) Seston C:P (molar)	N excretion rate per size class (μg N/g fish) P excretion rate (per size class) (μg P/g fish) Primary production rate (mg C/m^2/day) Seston C:N (molar) Seston C:P (molar)

Response

EM ID em.detail.idHelp ?	EM-668	Gizzard shad nutrient cycling: Burr Oak Lake	Gizzard shad nutrient cycling: Piedmont Lake	Gizzard shad nutrient cycling: Pleasant Hill Lake	Gizzard shad nutrient cycling: Kiser Lake	Gizzard shad nutrient cycling: O'Shaughnessy Lake	Gizzard shad nutrient cycling: C.J. Brown Lake	Gizzard shad nutrient cycling:Acton Lake
Observed Response Variables (and Units) em.detail.observedResponseHelp ?	None	*	*	*	*	*	*	*
Computed Response Variables (and Units) em.detail.computedResponseHelp ?	Lake trophic status (Not applicable) Lake-wide N excretion rate (gizzard shad population) (mg N/m^2/day) Lake-wide P excretion rate (gizzard shad population) (mg P/m^2/day) Nutrient demand (P) (phytoplankton) (mg P/m^2/day) Nutrient demand (phytoplankton) (mg N or P/m^2/day) Proportion of primary productivity supported by gizzard shad (unitless (0-1)) Proportion of watershed land use in agriculture (%)	Lake-wide N excretion rate (gizzard shad population) (mg N/m^2/day) Lake-wide P excretion rate (gizzard shad population) (mg P/m^2/day) Nitrogen demand (phytoplankton) (mg N /m^2/day) Phosphorus demand (phytoplankton) (mg P/m^2/day) Proportion of primary productivity supported by gizzard shad (unitless (0-1)) Proportion of watershed land use in agriculture (%)	Lake-wide N excretion rate (gizzard shad population) (mg N/m^2/day) Lake-wide P excretion rate (gizzard shad population) (mg P/m^2/day) Nitrogen demand (phytoplankton) (mg N /m^2/day) Phosphorus demand (phytoplankton) (mg P/m^2/day) Proportion of primary productivity supported by gizzard shad (unitless (0-1)) Proportion of watershed land use in agriculture (%)	Lake-wide N excretion rate (gizzard shad population) (mg N/m^2/day) Lake-wide P excretion rate (gizzard shad population) (mg P/m^2/day) Nitrogen demand (phytoplankton) (mg N /m^2/day) Phosphorus demand (phytoplankton) (mg P/m^2/day) Proportion of primary productivity supported by gizzard shad (unitless (0-1)) Proportion of watershed land use in agriculture (%)	Lake-wide N excretion rate (gizzard shad population) (mg N/m^2/day) Lake-wide P excretion rate (gizzard shad population) (mg P/m^2/day) Nitrogen demand (phytoplankton) (mg N /m^2/day) Phosphorus demand (phytoplankton) (mg P/m^2/day) Proportion of primary productivity supported by gizzard shad (unitless (0-1)) Proportion of watershed land use in agriculture (%)	Lake-wide N excretion rate (gizzard shad population) (mg N/m^2/day) Lake-wide P excretion rate (gizzard shad population) (mg P/m^2/day) Nitrogen demand (phytoplankton) (mg N /m^2/day) Phosphorus demand (phytoplankton) (mg P/m^2/day) Proportion of primary productivity supported by gizzard shad (unitless (0-1)) Proportion of watershed land use in agriculture (%)	Lake-wide N excretion rate (gizzard shad population) (mg N/m^2/day) Lake-wide P excretion rate (gizzard shad population) (mg P/m^2/day) Nitrogen demand (phytoplankton) (mg N /m^2/day) Phosphorus demand (phytoplankton) (mg P/m^2/day) Proportion of primary productivity supported by gizzard shad (unitless (0-1)) Proportion of watershed land use in agriculture (%)	Lake-wide N excretion rate (gizzard shad population) (mg N/m^2/day) Lake-wide P excretion rate (gizzard shad population) (mg P/m^2/day) Nitrogen demand (phytoplankton) (mg N /m^2/day) Phosphorus demand (phytoplankton) (mg P/m^2/day) Proportion of primary productivity supported by gizzard shad (unitless (0-1)) Proportion of watershed land use in agriculture (%)

US EPA

EcoService Models Library (ESML)

View Runs

: Nutrient Cycling by gizzard shad, Ohio, USA (EM-668)

EM Identity and Description

Related EMs (for example, other versions or derivations of this EM) described in ESML

EM Modeling Approach

Comment:

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EM Locations, Environments, Ecology

Terrestrial location (Classification hierarchy: Continent > Country > U.S. State [United States only])

Marine location (Classification hierarchy: Realm > Region > Province > Ecoregion)

Centroid Lat/Long (Decimal Degree)

Scale of differentiation of organisms modeled

Taxonomic level and name of organisms or groups identified

EnviroAtlas URL

EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system

* Note that run information is shown only where run data differ from the "parent" entry shown at left

CICES v 4.3 - Common International Classification of Ecosystem Services (Section > Division > Group > Class)

National Ecosystem Services Classification System (NESCS) Plus

(Environmental Subclass > Ecological End-Product (EEP) > EEP Subclass > EEP Modifier)

EM Variable Names (and Units)

* Note that for runs, variable name is displayed only where data for that variable differed by run AND those differences were reported in the source document. Where differences occurred but were not reported, the variable is not listed. Click on variable name to view details.

Predictor

Intermediate

Response