EcoService Models Library (ESML)
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Compare EMs
Which comparison is best for me?EM Variables by Variable Role
One quick way to compare ecological models (EMs) is by comparing their variables. Predictor variables show what kinds of influences a model is able to account for, and what kinds of data it requires. Response variables show what information a model is capable of estimating.
This first comparison shows the names (and units) of each EM’s variables, side-by-side, sorted by variable role. Variable roles in ESML are as follows:
- Predictor Variables
- Time- or Space-Varying Variables
- Constants and Parameters
- Intermediate (Computed) Variables
- Response Variables
- Computed Response Variables
- Measured Response Variables
EM Variables by Category
A second way to use variables to compare EMs is by focusing on the kind of information each variable represents. The top-level categories in the ESML Variable Classification Hierarchy are as follows:
- Policy Regarding Use or Management of Ecosystem Resources
- Land Surface (or Water Body Bed) Cover, Use or Substrate
- Human Demographic Data
- Human-Produced Stressor or Enhancer of Ecosystem Goods and Services Production
- Ecosystem Attributes and Potential Supply of Ecosystem Goods and Services
- Non-monetary Indicators of Human Demand, Use or Benefit of Ecosystem Goods and Services
- Monetary Values
Besides understanding model similarities, sorting the variables for each EM by these 7 categories makes it easier to see if the compared models can be linked using similar variables. For example, if one model estimates an ecosystem attribute (in Category 5), such as water clarity, as a response variable, and a second model uses a similar attribute (also in Category 5) as a predictor of recreational use, the two models can potentially be used in tandem. This comparison makes it easier to spot potential model linkages.
All EM Descriptors
This selection allows a more detailed comparison of EMs by model characteristics other than their variables. The 50-or-so EM descriptors for each model are presented, side-by-side, in the following categories:
- EM Identity and Description
- EM Modeling Approach
- EM Locations, Environments, Ecology
- EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
EM Descriptors by Modeling Concepts
This feature guides the user through the use of the following seven concepts for comparing and selecting EMs:
- Conceptual Model
- Modeling Objective
- Modeling Context
- Potential for Model Linkage
- Feasibility of Model Use
- Model Certainty
- Model Structural Information
Though presented separately, these concepts are interdependent, and information presented under one concept may have relevance to other concepts as well.
EM Identity and Description
EM ID
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EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
EM Short Name
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InVEST (v1.004) Carbon, Indonesia | Nitrogen fixation rates, Guánica Bay, Puerto Rico | C sequestration in grassland restoration, England | WESP: Marsh & wet meadow, ID, USA |
EM Full Name
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InVEST (Integrated Valuation of Environmental Services and Tradeoffs version 1.004) carbon storage and sequestration, Sumatra, Indonesia | Nitrogen fixation rates, Guánica Bay, Puerto Rico, USA | Carbon sequestration in grassland diversity restoration, England | WESP: Seasonally flooded marsh & wet meadow, Idaho, USA |
EM Source or Collection
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InVEST | US EPA | None | None |
EM Source Document ID
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309 |
338 ?Comment:WE received a draft copy prior to journal publication that was agency reviewed. |
396 |
393 ?Comment:Additional data came from electronic appendix provided by author Chris Murphy. |
Document Author
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Bhagabati, N. K., Ricketts, T., Sulistyawan, T. B. S., Conte, M., Ennaanay, D., Hadian, O., McKenzie, E., Olwero, N., Rosenthal, A., Tallis, H., and Wolney, S. | Amelia Smith, Susan Harrell Yee, Marc Russell, Jill Awkerman and William S. Fisher | De Deyn, G. B., R. S. Shiel, N. J. Ostle, N. P. McNamara, S. Oakley, I. Young, C. Freeman, N. Fenner, H. Quirk, and R. D. Bardgett | Murphy, C. and T. Weekley |
Document Year
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2014 | 2017 | 2011 | 2012 |
Document Title
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Ecosystem services reinforce Sumatran tiger conservation in land use plans | Linking ecosystem services supply to stakeholder concerns on both land and sea: An example from Guanica Bay watershed, Puerto Rico | Additional carbon sequestration benefits of grassland diversity restoration | Measuring outcomes of wetland restoration, enhancement, and creation in Idaho-- Assessing potential functions, values, and condition in a watershed context. |
Document Status
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Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published |
Comments on Status
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Published journal manuscript | Published journal manuscript | Published journal manuscript | Published report |
EM ID
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EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
https://www.naturalcapitalproject.org/invest/ | Not applicable | Not applicable | Not applicable | |
Contact Name
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Nirmal K. Bhagabati | Susan H. Yee | Gerlinde B. De Deyn | Chris Murphy |
Contact Address
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The Nature Conservancy, 1107 Laurel Avenue, Felton, CA 95018 | U.S. Environmental Protection Agency, Gulf Ecology Division, Gulf Breeze, FL 32561, USA | Dept. of Terrestrial Ecology, Netherlands Institute of Ecology, P O Box 40, 6666 ZG Heteren, The Netherlands | Idaho Dept. Fish and Game, Wildlife Bureau, Habitat Section, Boise, ID |
Contact Email
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nirmal.bhagabati@wwfus.org | yee.susan@epa.gov | g.dedeyn@nioo.knaw.nl; gerlindede@gmail.com | chris.murphy@idfg.idaho.gov |
EM ID
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EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
Summary Description
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Please note: This ESML entry describes a specific, published application of an InVEST model. Different versions (e.g. different tiers) or more recent versions of this model may be available at the InVEST website. ABSTRACT: "...Here we use simple spatial analyses on readily available datasets to compare the distribution of five ecosystem services with tiger habitat in central Sumatra. We assessed services and habitat in 2008 and the changes in these variables under two future scenarios: a conservation-friendly Green Vision, and a Spatial Plan developed by the Indonesian government..." AUTHOR'S DESCRIPTION: "We used a modeling tool, InVEST (Integrated Valuation of Environmental Services and Tradeoffs version 1.004; Tallis et al., 2010), to map and quantify tiger habitat quality and five ecosystem services. InVEST maps ecosystem services and the quality of species habitat as production functions of LULC using simple biophysical models. Models were parameterized using data from regional agencies, literature surveys, global databases, site visits and prior field experience (Table 1)... We mapped biomass carbon by assigning carbon values (in ton ha_1) for aboveground, belowground, and dead organic matter to each LULC class based on values from literature, as described in Tallis et al. (2010). We mapped soil carbon separately, as large quantities of carbon are stored in peat soil (Page et al., 2011). We estimated total losses in peat carbon over 50 years into the future scenarios, using reported annual emission rates for specific LULC transitions on peat (Uryu et al., 2008)...Although InVEST reports ecosystem services in biophysical units, its simple models are best suited to understanding broad patterns of spatial variation (Tallis and Polasky, 2011), rather than for precise quantification. Additionally, we lacked field measurements against which to calibrate our outputs. Therefore, we focused on relative spatial distribution across the landscape, and relative change to scenarios." | AUTHOR'S DESCRIPTION: " …In Guánica Bay watershed, Puerto Rico, deforestation and drainage of a large lagoon have led to sediment, contaminant, and nutrient transport into the bay, resulting in declining quality of coral reefs. A watershed management plan is currently being implemented to restore reefs through a variety of proposed actions…After the workshops, fifteen indicators of terrestrial ecosystem services in the watershed and four indicators in the coastal zone were identified to reflect the wide range of stakeholder concerns that could be impacted by management decisions. Ecosystem service production functions were applied to quantify and map ecosystem services supply in the Guánica Bay watershed, as well as an additional highly engineered upper multi-watershed area connected to the lower watershed via a series of reservoirs and tunnels,…” AUTHOR''S DESCRIPTION: "The U.S. Coral Reef Task Force (CRTF), a collaboration of federal, state and territorial agencies, initiated a program in 2009 to better incorporate land-based sources of pollution and socio-economic considerations into watershed strategies for coral reef protection (Bradley et al., 2016)...Baseline measures for relevant ecosystem services were calculated by parameterizing existing methods, largely based on land cover (Egoh et al., 2012; Martinez- Harms and Balvanera, 2012), with relevant rates of ecosystem services production for Puerto Rico, and applying them to map ecosystem services supply for the Guánica Bay Watershed...The Guánica Bay watershed is a highly engineered watershed in southwestern Puerto Rico, with a series of five reservoirs and extensive tunnel systems artificially connecting multiple mountainous sub-watersheds to the lower watershed of the Rio Loco, which itself is altered by an irrigation canal and return drainage ditch that diverts water through the Lajas Valley (PRWRA, 1948)...For each objective, a translator of ecosystem services production, i.e., ecological production function, was used to quantify baseline measurements of ecosystem services supply from land use/land cover (LULC) maps for watersheds across Puerto Rico...Two additional metrics, nitrogen fixation and rates of carbon sequestration into soil and sediment, were also calculated as potential measures of soil quality and agricultural productivity. Carbon sequestration and nitrogen fixation rates were assigned to each land cover class" | ABSTRACT: "A major aim of European agri-environment policy is the management of grassland for botanical diversity conservation and restoration, together with the delivery of ecosystem services including soil carbon (C) sequestration. To test whether management for biodiversity restoration has additional benefits for soil C sequestration, we investigated C and nitrogen (N) accumulation rates in soil and C and N pools in vegetation in a long-term field experiment (16 years) in which fertilizer application and plant seeding were manipulated. In addition, the abundance of the legume Trifolium pratense was manipulated for the last 2 years. To unravel the mechanisms underlying changes in soil C and N pools, we also tested for effects of diversity restoration management on soil structure, ecosystem respiration and soil enzyme activities…" AUTHOR'S DESCRIPTION: "Measurements were made on 36 plots of 3 x 3 m comprising two management treatments (and their controls) in a long-term multifactorial grassland restoration experiment which have successfully increased plant species diversity, namely the cessation of NPK fertilizer application and the addition of seed mixtures…" | A wetland restoration monitoring and assessment program framework was developed for Idaho. The project goal was to assess outcomes of substantial governmental and private investment in wetland restoration, enhancement and creation. The functions, values, condition, and vegetation at restored, enhanced, and created wetlands on private and state lands across Idaho were retrospectively evaluated. Assessment was conducted at multiple spatial scales and intensities. Potential functions and values (ecosystem services) were rapidly assessed using the Oregon Rapid Wetland Assessment Protocol. Vegetation samples were analyzed using Floristic Quality Assessment indices from Washington State. We compared vegetation of restored, enhanced, and created wetlands with reference wetlands that occurred in similar hydrogeomorphic environments determined at the HUC 12 level. |
Specific Policy or Decision Context Cited
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This analysis provided input to government-led spatial planning and strategic environmental assessments in the study area. This region contains some of the last remaining forest habitat of the critically endangered Sumatran tiger, Panthera tigris sumatrae. | None provided | None identified | None identified |
Biophysical Context
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Six watersheds in central Sumatra covering portions of Riau, Jambi and West Sumatra provinces. The Barisan mountain range comprises the western edge of the watersheds, while peat swamps predominate in the east. | No additional description provided | Lolium perenne-Cynosorus cristatus grassland; The soil is a shallow brown-earth (average depth 28 cm) over limestone of moderate-high residual fertility. | restored, enhanced and created wetlands |
EM Scenario Drivers
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Baseline year 2008, future LULC Sumatra 2020 Roadmap (Vision), future LULC Government Spatial Plan | No scenarios presented | Additional benefits due to biodiversity restoration practices | Sites, function or habitat focus |
EM ID
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EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
Method Only, Application of Method or Model Run
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Method + Application (multiple runs exist) View EM Runs | Method + Application | Method + Application (multiple runs exist) View EM Runs | Method + Application (multiple runs exist) View EM Runs |
New or Pre-existing EM?
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Application of existing model | Application of existing model | New or revised model | New or revised model |
Related EMs (for example, other versions or derivations of this EM) described in ESML
EM ID
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EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
Document ID for related EM
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Doc-315 | None | None | Doc-390 |
EM ID for related EM
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EM-374 | None | None | EM-718 | EM-734 | EM-743 |
EM Modeling Approach
EM ID
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EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
EM Temporal Extent
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2008-2020 | 1978 - 2009 | 1990-2007 | 2010-2012 |
EM Time Dependence
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time-stationary | time-stationary | time-stationary | time-dependent |
EM Time Reference (Future/Past)
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Not applicable | Not applicable | Not applicable | past time |
EM Time Continuity
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Not applicable | Not applicable | Not applicable | Not applicable |
EM Temporal Grain Size Value
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Not applicable | Not applicable | Not applicable | Not applicable |
EM Temporal Grain Size Unit
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Not applicable | Not applicable | Not applicable | Not applicable |
EM ID
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EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
Bounding Type
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Watershed/Catchment/HUC | Watershed/Catchment/HUC | Other | Multiple unrelated locations (e.g., meta-analysis) |
Spatial Extent Name
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central Sumatra | Guanica Bay watershed | Colt Park meadows, Ingleborough National Nature Reserve, northern England | Wetlands in idaho |
Spatial Extent Area (Magnitude)
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100,000-1,000,000 km^2 | 100-1000 km^2 | <1 ha | 100,000-1,000,000 km^2 |
EM ID
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EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
EM Spatial Distribution
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spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially lumped (in all cases) |
Spatial Grain Type
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area, for pixel or radial feature | other (specify), for irregular (e.g., stream reach, lake basin) | area, for pixel or radial feature | Not applicable |
Spatial Grain Size
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30 m x 30 m | HUC | 3 m x 3 m | Not applicable |
EM ID
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EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
EM Computational Approach
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Analytic | Analytic | Analytic | Numeric |
EM Determinism
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deterministic | deterministic | stochastic | deterministic |
Statistical Estimation of EM
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EM ID
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EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
Model Calibration Reported?
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No | No | Not applicable | No |
Model Goodness of Fit Reported?
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No | No | Not applicable | No |
Goodness of Fit (metric| value | unit)
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None | None | None | None |
Model Operational Validation Reported?
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No | No | No | No |
Model Uncertainty Analysis Reported?
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No | No | No | No |
Model Sensitivity Analysis Reported?
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No | No | No | No |
Model Sensitivity Analysis Include Interactions?
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Not applicable | Not applicable | Not applicable | Not applicable |
EM Locations, Environments, Ecology
Terrestrial location (Classification hierarchy: Continent > Country > U.S. State [United States only])
EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
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Marine location (Classification hierarchy: Realm > Region > Province > Ecoregion)
EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
None | None | None | None |
Centroid Lat/Long (Decimal Degree)
EM ID
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EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
Centroid Latitude
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0 | 17.96 | 54.2 | 44.06 |
Centroid Longitude
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102 | -67.02 | -2.35 | -114.69 |
Centroid Datum
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WGS84 | WGS84 | WGS84 | WGS84 |
Centroid Coordinates Status
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Provided | Estimated | Provided | Estimated |
EM ID
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EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
EM Environmental Sub-Class
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Inland Wetlands | Lakes and Ponds | Forests | Agroecosystems | Created Greenspace | Grasslands | Scrubland/Shrubland | Barren | Inland Wetlands | Forests | Agroecosystems | Created Greenspace | Grasslands | Scrubland/Shrubland | Barren | Agroecosystems | Grasslands | Inland Wetlands |
Specific Environment Type
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104 land use land cover classes | Tropical terrestrial | fertilized grassland (historically hayed) | created, restored and enhanced wetlands |
EM Ecological Scale
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Ecological scale is finer than that of the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class |
Scale of differentiation of organisms modeled
EM ID
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EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
EM Organismal Scale
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Community | Not applicable | Community | Not applicable |
Taxonomic level and name of organisms or groups identified
EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
None Available | None Available | None Available | None Available |
EnviroAtlas URL
EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
CICES v 4.3 - Common International Classification of Ecosystem Services (Section > Division > Group > Class)
EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
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<a target="_blank" rel="noopener noreferrer" href="https://www.epa.gov/eco-research/national-ecosystem-services-classification-system-nescs-plus">National Ecosystem Services Classification System (NESCS) Plus</a>
(Environmental Subclass > Ecological End-Product (EEP) > EEP Subclass > EEP Modifier)
EM-349 ![]() |
EM-432 |
EM-735 ![]() |
EM-760 ![]() |
None |
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None | None |