EcoService Models Library (ESML)
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Compare EMs
Which comparison is best for me?EM Variables by Variable Role
One quick way to compare ecological models (EMs) is by comparing their variables. Predictor variables show what kinds of influences a model is able to account for, and what kinds of data it requires. Response variables show what information a model is capable of estimating.
This first comparison shows the names (and units) of each EM’s variables, side-by-side, sorted by variable role. Variable roles in ESML are as follows:
- Predictor Variables
- Time- or Space-Varying Variables
- Constants and Parameters
- Intermediate (Computed) Variables
- Response Variables
- Computed Response Variables
- Measured Response Variables
EM Variables by Category
A second way to use variables to compare EMs is by focusing on the kind of information each variable represents. The top-level categories in the ESML Variable Classification Hierarchy are as follows:
- Policy Regarding Use or Management of Ecosystem Resources
- Land Surface (or Water Body Bed) Cover, Use or Substrate
- Human Demographic Data
- Human-Produced Stressor or Enhancer of Ecosystem Goods and Services Production
- Ecosystem Attributes and Potential Supply of Ecosystem Goods and Services
- Non-monetary Indicators of Human Demand, Use or Benefit of Ecosystem Goods and Services
- Monetary Values
Besides understanding model similarities, sorting the variables for each EM by these 7 categories makes it easier to see if the compared models can be linked using similar variables. For example, if one model estimates an ecosystem attribute (in Category 5), such as water clarity, as a response variable, and a second model uses a similar attribute (also in Category 5) as a predictor of recreational use, the two models can potentially be used in tandem. This comparison makes it easier to spot potential model linkages.
All EM Descriptors
This selection allows a more detailed comparison of EMs by model characteristics other than their variables. The 50-or-so EM descriptors for each model are presented, side-by-side, in the following categories:
- EM Identity and Description
- EM Modeling Approach
- EM Locations, Environments, Ecology
- EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
EM Descriptors by Modeling Concepts
This feature guides the user through the use of the following seven concepts for comparing and selecting EMs:
- Conceptual Model
- Modeling Objective
- Modeling Context
- Potential for Model Linkage
- Feasibility of Model Use
- Model Certainty
- Model Structural Information
Though presented separately, these concepts are interdependent, and information presented under one concept may have relevance to other concepts as well.
EM Identity and Description
EM ID
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EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
EM Short Name
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Green biomass production, Central French Alps | Community flowering date, Central French Alps | PATCH, western USA | Value of a reef dive site, St. Croix, USVI | Seed mix for native plant establishment, IA, USA |
EM Full Name
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Green biomass production, Central French Alps | Community weighted mean flowering date, Central French Alps | PATCH (Program to Assist in Tracking Critical Habitat), western USA | Value of a dive site (reef), St. Croix, USVI | Cost-effective seed mix design for native plant establishment, Iowa, USA |
EM Source or Collection
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EU Biodiversity Action 5 | EU Biodiversity Action 5 | US EPA | US EPA | None |
EM Source Document ID
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260 | 260 | 2 | 335 | 394 |
Document Author
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Lavorel, S., Grigulis, K., Lamarque, P., Colace, M-P, Garden, D., Girel, J., Pellet, G., and Douzet, R. | Lavorel, S., Grigulis, K., Lamarque, P., Colace, M-P, Garden, D., Girel, J., Pellet, G., and Douzet, R. | Carroll, C, Phillips, M. K. , Lopez-Gonzales, C. A and Schumaker, N. H. | Yee, S. H., Dittmar, J. A., and L. M. Oliver | Meissen, J. |
Document Year
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2011 | 2011 | 2006 | 2014 | 2018 |
Document Title
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Using plant functional traits to understand the landscape distribution of multiple ecosystem services | Using plant functional traits to understand the landscape distribution of multiple ecosystem services | Defining recovery goals and strategies for endangered species: The wolf as a case study | Comparison of methods for quantifying reef ecosystem services: A case study mapping services for St. Croix, USVI | Cost-effective seed mix design and first-year management |
Document Status
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Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published |
Comments on Status
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Published journal manuscript | Published journal manuscript | Published journal manuscript | Published journal manuscript | Published report |
EM ID
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EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | |
Contact Name
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Sandra Lavorel | Sandra Lavorel | Carlos Carroll | Susan H. Yee | Justin Meissen |
Contact Address
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Laboratoire d’Ecologie Alpine, UMR 5553 CNRS Université Joseph Fourier, BP 53, 38041 Grenoble Cedex 9, France | Laboratoire d’Ecologie Alpine, UMR 5553 CNRS Université Joseph Fourier, BP 53, 38041 Grenoble Cedex 9, France | Klamath Center for Conservation Research, Orleans, CA 95556 | US EPA, Office of Research and Development, NHEERL, Gulf Ecology Division, Gulf Breeze, FL 32561, USA | Tallgrass Prairie Center, University of Northern Iowa |
Contact Email
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sandra.lavorel@ujf-grenoble.fr | sandra.lavorel@ujf-grenoble.fr | carlos@cklamathconservation.org | yee.susan@epa.gov | Not reported |
EM ID
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EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
Summary Description
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ABSTRACT: "Here, we propose a new approach for the analysis, mapping and understanding of multiple ES delivery in landscapes. Spatially explicit single ES models based on plant traits and abiotic characteristics are combined to identify ‘hot’ and ‘cold’ spots of multiple ES delivery, and the land use and biotic determinants of such distributions. We demonstrate the value of this trait-based approach as compared to a pure land-use approach for a pastoral landscape from the central French Alps, and highlight how it improves understanding of ecological constraints to, and opportunities for, the delivery of multiple services. Vegetative height and leaf traits such as leaf dry matter content were response traits strongly influenced by land use and abiotic environment, with follow-on effects on several ecosystem properties (e.g., green biomass production), and could therefore be used as functional markers of ES." AUTHOR'S DESCRIPTION: "Variation in green biomass production was modelled using…traits community-weighted mean (CWM) and functional divergence (FD) and abiotic variables (continuous variables; trait + abiotic) following Diaz et al. (2007). …The comparison between this model and the land-use alone model identifies the need for site-based information beyond a land use or land cover proxy, and the comparison with the land use + abiotic model assesses the value of additional ecological (trait) information…Green biomass production for each pixel was calculated and mapped using model estimates for…regression coefficients on abiotic variables and traits. For each pixel these calculations were applied to mapped estimates of abiotic variables and trait CWM and FD. This step is critically novel as compared to a direct application of the model by Diaz et al. (2007) in that we explicitly modelled the responses of trait community-weighted means and functional divergences to environment prior to evaluating their effects on ecosystem properties. Such an approach is the key to the explicit representation of functional variation across the landscape, as opposed to the use of unique trait values within each land use (see Albert et al. 2010)." | ABSTRACT: "Here, we propose a new approach for the analysis, mapping and understanding of multiple ES delivery in landscapes. Spatially explicit single ES models based on plant traits and abiotic characteristics are combined to identify ‘hot’ and ‘cold’ spots of multiple ES delivery, and the land use and biotic determinants of such distributions. We demonstrate the value of this trait-based approach as compared to a pure land-use approach for a pastoral landscape from the central French Alps, and highlight how it improves understanding of ecological constraints to, and opportunities for, the delivery of multiple services." AUTHOR'S DESCRIPTION: "Community-weighted mean date of flowering onset was modelled using mixed models with land use and abiotic variables as fixed effects (LU + abiotic model) and year as a random effect…and modelled for each 20 x 20 m pixel using GLM estimated effects for each land use category and estimated regression coefficients with abiotic variables." | **Note: A more recent version of this model exists. See Related EMs below for links to related models/applications.** AUTHORS' DESCRIPTION: "PATCH (program to assist in tracking critical habitat), the SEPM used here, is designed for studying territorial vertebrates. It links the survival and fecundity of individual animals to geographic information system (GIS) data on mortality risk and habitat productivity at the scale of an individual or pack territory. Territories are allocated by intersecting the GIS data with an array of hexagonal cells. The different habitat types in the GIS maps are assigned weights based on the relative levels of fecundity and survival expected in those habitat classes. Base survival and reproductive rates, derived from published field studies, are then supplied to the model as a population projection matrix. The model scales these base matrix values using the mean of the habitat weights within each hexagon, with lower means translating into lower survival rates or reproductive output. Each individual in the population is tracked through a yearly cycle of survival, fecundity, and dispersal events. Environmental stochasticity is incorporated by drawing each year’s base population matrix from a randomized set of matrices whose elements were drawn from a beta (survival) or normal (fecundity) distribution. Adult organisms are classified as either territorial or floaters. The movement of territorial individuals is governed by a parameter for site fidelity, but floaters must always search for available breeding sites. As pack size increases, pack members in the model have a greater tendency to disperse and search for new available breeding sites. Movement decisions use a directed random walk that combines varying proportions of randomness, correlation, and attraction to higher-quality habitat (Schumaker 1998)." | ABSTRACT: "...We investigated and compared a number of existing methods for quantifying ecological integrity, shoreline protection, recreational opportunities, fisheries production, and the potential for natural products discovery from reefs. Methods were applied to mapping potential ecosystem services production around St. Croix, U.S. Virgin Islands. Overall, we found that a number of different methods produced similar predictions." AUTHOR'S DESCRIPTION: "A number of methods have been developed for linking biophysical attributes of reef condition, such as reef structural complexity, fish biomass, or species richness, to provisioning of ecosystem goods and services (Principe et al., 2012). We investigated the feasibility of using existing methods and data for mapping production of reef ecosystem goods and services. We applied these methods toward mapping potential ecosystem goods and services production in St. Croix, U.S. Virgin Islands (USVI)...For each of the five categories of ecosystem services, we chose a suite of models and indices for estimating potential production based on relative ease of implementation, consisting of well-defined parameters, and likely availability of input data, to maximize potential for transferability to other locations. For each method, we assembled the necessary reef condition and environmental data as spatial data layers for St. Croix (Table1). The coastal zone surrounding St. Croix was divided into 10x10 m grid cells, and production functions were applied to quantify ecosystem services provisioning in each grid cell...A number of recreational activities are associated directly or indirectly with coral reefs including scuba diving, snorkeling, surfing, underwater photography, recreational fishing, wildlife viewing, beach sunbathing and swimming, and beachcombing (Principe et al., 2012)…Another method to quantify recreational opportunities is to use survey data of tourists and recreational visitors to the reefs to generate statistical models to quantify the link between reef condition and production of recreation-related ecosystem services. Wielgus et al. (2003) used interviews with SCUBA divers in Israel to derive coefficients for a choice model in which willingness to pay for higher quality dive sites was determined in part by a weighted combination of factors identified with dive quality: Relative value of dive site = 0.1227(Scoral+Sfish+Acoral+Afish)+0.0565V where Scoral, Sfish are coral and fish richness, Acoral, Afish are abundances of fish and coral per square meter, and V is water visibility (meters)." | AUTHOR'S DESCRIPTION: "Restoring ecosystem services at scale requires executing conservation programs in a way that is resource and cost efficient as well as ecologically effective…Seed mix design is one of the largest determinants of project cost and ecological outcomes for prairie reconstructions. In particular, grass-to-forb seeding ratio affects cost since forb seed can be much more expensive relative to grass species (Prairie Moon Nursery 2012). Even for seed mixes with the same overall seeding rates, a mix with a low grass-to-forb seeding ratio is considerably more expensive than one with a high grass-to-forb ratio. Seeding rates for different plant functional groups that are too high or low may also adversely affect ecological outcomes…First-year management may also play a role in cost-effective prairie reconstruction. Post-agricultural sites where restoration typically occurs are often quickly dominated by fast-growing annual weeds by the time sown prairie seeds begin germinating (Smith et al. 2010)… Williams and others (2007) showed that prairie seedlings sown into established warm-season grasses were reliant on high light conditions created by frequently mowing tall vegetation in order to survive in subsequent years…Our objective was to compare native plant establishment and cost effectiveness with and without first-year mowing for three different seed mixes that differed in grass to forb ratio and soil type customization. With knowledge of plant establishment, cost effectiveness, and mowing management outcomes, conservation practitioners will be better equipped to restore prairie efficiently and successfully." |
Specific Policy or Decision Context Cited
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None identified | None identified | AUTHOR DESCRIPTION: "Comprehensive habitat and viability assessments. . . [more rigoursly defined] can clarify debate of goals for recovery of large carnivores"; Endangered Species Act and related litigation | None identified | Seed mix design and management practices for native plant restoration |
Biophysical Context
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Elevation ranges from 1552 to 2442 m, on predominately south-facing slopes | Elevation ranges from 1552 to 2442 m, on predominantly south-facing slopes | Great Plains to Pacific Coast, northern Rocky Mountains, Pacific Northwest | No additional description provided | The soils underlying the study site are primarily poorly drained Clyde clay loams, with a minor component of somewhat poorly drained Floyd loams in the northwest (NRCS 2016). Topographically, the study site is level, and slopes do not exceed 5% grade. Land use prior to this experiment was agricultural, with corn and soybeans consistently grown in rotation at the site. |
EM Scenario Drivers
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No scenarios presented | No scenarios presented | Population growth, road development (density) on public vs private land | No scenarios presented | No scenarios presented |
EM ID
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EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
Method Only, Application of Method or Model Run
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Method + Application | Method + Application | Method + Application (multiple runs exist) View EM Runs | Method + Application | Method + Application (multiple runs exist) View EM Runs |
New or Pre-existing EM?
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New or revised model | New or revised model | New or revised model | Application of existing model | New or revised model |
Related EMs (for example, other versions or derivations of this EM) described in ESML
EM ID
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EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
Document ID for related EM
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Doc-260 | Doc-260 | Doc-269 | Doc-328 | Doc-337 | None | Doc-395 |
EM ID for related EM
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EM-66 | EM-68 | EM-69 | EM-70 | EM-71 | EM-79 | EM-80 | EM-81 | EM-82 | EM-83 | EM-65 | EM-66 | EM-68 | EM-69 | EM-70 | EM-79 | EM-80 | EM-81 | EM-82 | EM-83 | EM-403 | EM-422 | None | EM-728 |
EM Modeling Approach
EM ID
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EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
EM Temporal Extent
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2007-2009 | 2007-2008 | 2000-2025 | 2006-2007, 2010 | 2015-2017 |
EM Time Dependence
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time-stationary | time-stationary | time-dependent | time-stationary | time-dependent |
EM Time Reference (Future/Past)
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Not applicable | Not applicable | future time | Not applicable | Not applicable |
EM Time Continuity
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Not applicable | Not applicable | discrete | Not applicable | discrete |
EM Temporal Grain Size Value
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Not applicable | Not applicable | 1 | Not applicable | 1 |
EM Temporal Grain Size Unit
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Not applicable | Not applicable | Year | Not applicable | Year |
EM ID
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EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
Bounding Type
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Physiographic or Ecological | Physiographic or Ecological | Physiographic or ecological | Physiographic or ecological | Other |
Spatial Extent Name
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Central French Alps | Central French Alps | Western United States | Coastal zone surrounding St. Croix | Iowa State University Northeast Research and Demonstration Farm |
Spatial Extent Area (Magnitude)
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10-100 km^2 | 10-100 km^2 | >1,000,000 km^2 | 100-1000 km^2 | <1 ha |
EM ID
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EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
EM Spatial Distribution
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spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially distributed (in at least some cases) |
Spatial Grain Type
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area, for pixel or radial feature | area, for pixel or radial feature | area, for pixel or radial feature | area, for pixel or radial feature | area, for pixel or radial feature |
Spatial Grain Size
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20 m x 20 m | 20 m x 20 m | 504 km^2 | 10 m x 10 m | 20 ft x 28 ft |
EM ID
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EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
EM Computational Approach
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Analytic | Analytic | Numeric | Analytic | Analytic |
EM Determinism
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deterministic | deterministic | stochastic | deterministic | stochastic |
Statistical Estimation of EM
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EM ID
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EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
Model Calibration Reported?
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No | No | Unclear | Yes | Not applicable |
Model Goodness of Fit Reported?
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Yes | Yes | No | No | Not applicable |
Goodness of Fit (metric| value | unit)
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None | None | None |
Model Operational Validation Reported?
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Yes | No | No | Yes | No |
Model Uncertainty Analysis Reported?
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No | No | No | No | Not applicable |
Model Sensitivity Analysis Reported?
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No | No |
Yes ?Comment:No results reported. Just a general statement was made about PATCH sensitivity and that demographic parameters are more sensitive that variation in other parameters such as dispersadistance . Reference made to another publication Carroll et al. 2003. Use of population viability analysis and reserve slelection algorithms in regional conservation plans. Ecol. App. 13:1773-1789. |
No | Not applicable |
Model Sensitivity Analysis Include Interactions?
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Not applicable | Not applicable | Unclear | Not applicable | Not applicable |
EM Locations, Environments, Ecology
Terrestrial location (Classification hierarchy: Continent > Country > U.S. State [United States only])
EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
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None |
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Marine location (Classification hierarchy: Realm > Region > Province > Ecoregion)
EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
None | None | None |
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None |
Centroid Lat/Long (Decimal Degree)
EM ID
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EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
Centroid Latitude
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45.05 | 45.05 | 39.88 | 17.73 | 42.93 |
Centroid Longitude
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6.4 | 6.4 | -113.81 | -64.77 | -92.57 |
Centroid Datum
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WGS84 | WGS84 | WGS84 | WGS84 | WGS84 |
Centroid Coordinates Status
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Provided | Provided | Estimated | Estimated | Provided |
EM ID
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EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
EM Environmental Sub-Class
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Agroecosystems | Grasslands | Agroecosystems | Grasslands | Terrestrial Environment (sub-classes not fully specified) | Near Coastal Marine and Estuarine | Agroecosystems | Grasslands |
Specific Environment Type
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Subalpine terraces, grasslands, and meadows | Subalpine terraces, grasslands, and meadows. | Not reported | Coral reefs | Research farm in historic grassland |
EM Ecological Scale
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Not applicable | Not applicable | Ecological scale corresponds to the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class |
Scale of differentiation of organisms modeled
EM ID
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EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
EM Organismal Scale
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Community | Community | Species | Guild or Assemblage | Community |
Taxonomic level and name of organisms or groups identified
EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
None Available | None Available |
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None Available | None Available |
EnviroAtlas URL
EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
GAP Ecological Systems | None Available | Dasymetric Allocation of Population | None Available | GAP Ecological Systems |
EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
CICES v 4.3 - Common International Classification of Ecosystem Services (Section > Division > Group > Class)
EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
None | None |
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<a target="_blank" rel="noopener noreferrer" href="https://www.epa.gov/eco-research/national-ecosystem-services-classification-system-nescs-plus">National Ecosystem Services Classification System (NESCS) Plus</a>
(Environmental Subclass > Ecological End-Product (EEP) > EEP Subclass > EEP Modifier)
EM-65 | EM-71 |
EM-98 ![]() |
EM-455 |
EM-719 ![]() |
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None |
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