EcoService Models Library (ESML)
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Compare EMs
Which comparison is best for me?EM Variables by Variable Role
One quick way to compare ecological models (EMs) is by comparing their variables. Predictor variables show what kinds of influences a model is able to account for, and what kinds of data it requires. Response variables show what information a model is capable of estimating.
This first comparison shows the names (and units) of each EM’s variables, side-by-side, sorted by variable role. Variable roles in ESML are as follows:
- Predictor Variables
- Time- or Space-Varying Variables
- Constants and Parameters
- Intermediate (Computed) Variables
- Response Variables
- Computed Response Variables
- Measured Response Variables
EM Variables by Category
A second way to use variables to compare EMs is by focusing on the kind of information each variable represents. The top-level categories in the ESML Variable Classification Hierarchy are as follows:
- Policy Regarding Use or Management of Ecosystem Resources
- Land Surface (or Water Body Bed) Cover, Use or Substrate
- Human Demographic Data
- Human-Produced Stressor or Enhancer of Ecosystem Goods and Services Production
- Ecosystem Attributes and Potential Supply of Ecosystem Goods and Services
- Non-monetary Indicators of Human Demand, Use or Benefit of Ecosystem Goods and Services
- Monetary Values
Besides understanding model similarities, sorting the variables for each EM by these 7 categories makes it easier to see if the compared models can be linked using similar variables. For example, if one model estimates an ecosystem attribute (in Category 5), such as water clarity, as a response variable, and a second model uses a similar attribute (also in Category 5) as a predictor of recreational use, the two models can potentially be used in tandem. This comparison makes it easier to spot potential model linkages.
All EM Descriptors
This selection allows a more detailed comparison of EMs by model characteristics other than their variables. The 50-or-so EM descriptors for each model are presented, side-by-side, in the following categories:
- EM Identity and Description
- EM Modeling Approach
- EM Locations, Environments, Ecology
- EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
EM Descriptors by Modeling Concepts
This feature guides the user through the use of the following seven concepts for comparing and selecting EMs:
- Conceptual Model
- Modeling Objective
- Modeling Context
- Potential for Model Linkage
- Feasibility of Model Use
- Model Certainty
- Model Structural Information
Though presented separately, these concepts are interdependent, and information presented under one concept may have relevance to other concepts as well.
EM Identity and Description
EM ID
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EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
EM Short Name
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Green biomass production, Central French Alps | Community flowering date, Central French Alps | Divergence in flowering date, Central French Alps | Annual profit - carbon plantings, South Australia | Biological pest control, Uppland Province, Sweden |
EM Full Name
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Green biomass production, Central French Alps | Community weighted mean flowering date, Central French Alps | Functional divergence in flowering date, Central French Alps | Annual profit from carbon plantings, South Australia | Biological control of agricultural pests by natural predators, Uppland Province, Sweden |
EM Source or Collection
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EU Biodiversity Action 5 | EU Biodiversity Action 5 | EU Biodiversity Action 5 | None | None |
EM Source Document ID
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260 | 260 | 260 | 243 | 299 |
Document Author
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Lavorel, S., Grigulis, K., Lamarque, P., Colace, M-P, Garden, D., Girel, J., Pellet, G., and Douzet, R. | Lavorel, S., Grigulis, K., Lamarque, P., Colace, M-P, Garden, D., Girel, J., Pellet, G., and Douzet, R. | Lavorel, S., Grigulis, K., Lamarque, P., Colace, M-P, Garden, D., Girel, J., Pellet, G., and Douzet, R. | Crossman, N. D., Bryan, B. A., and Summers, D. M. | Jonsson, M., Bommarco, R., Ekbom, B., Smith, H.G., Bengtsson, J., Caballero-Lopez, B., Winqvist, C., and Olsson, O. |
Document Year
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2011 | 2011 | 2011 | 2011 | 2014 |
Document Title
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Using plant functional traits to understand the landscape distribution of multiple ecosystem services | Using plant functional traits to understand the landscape distribution of multiple ecosystem services | Using plant functional traits to understand the landscape distribution of multiple ecosystem services | Carbon payments and low-cost conservation | Ecological production functions for biological control services in agricultural landscapes |
Document Status
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Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published |
Comments on Status
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Published journal manuscript | Published journal manuscript | Published journal manuscript | Published journal manuscript | Published journal manuscript |
EM ID
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EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | |
Contact Name
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Sandra Lavorel | Sandra Lavorel | Sandra Lavorel | Neville D. Crossman | Mattias Jonsson |
Contact Address
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Laboratoire d’Ecologie Alpine, UMR 5553 CNRS Université Joseph Fourier, BP 53, 38041 Grenoble Cedex 9, France | Laboratoire d’Ecologie Alpine, UMR 5553 CNRS Université Joseph Fourier, BP 53, 38041 Grenoble Cedex 9, France | Laboratoire d’Ecologie Alpine, UMR 5553 CNRS Université Joseph Fourier, BP 53, 38041 Grenoble Cedex 9, France | CSIRO Ecosystem Sciences, PMB 2, Glen Osmond, South Australia, 5064, Australia | Department of Ecology, Swedish University of Agricultural Sciences, PO Box 7044, SE-750 07 Uppsala, Sweden |
Contact Email
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sandra.lavorel@ujf-grenoble.fr | sandra.lavorel@ujf-grenoble.fr | sandra.lavorel@ujf-grenoble.fr | neville.crossman@csiro.au | mattias.jonsson@slu.se |
EM ID
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EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
Summary Description
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ABSTRACT: "Here, we propose a new approach for the analysis, mapping and understanding of multiple ES delivery in landscapes. Spatially explicit single ES models based on plant traits and abiotic characteristics are combined to identify ‘hot’ and ‘cold’ spots of multiple ES delivery, and the land use and biotic determinants of such distributions. We demonstrate the value of this trait-based approach as compared to a pure land-use approach for a pastoral landscape from the central French Alps, and highlight how it improves understanding of ecological constraints to, and opportunities for, the delivery of multiple services. Vegetative height and leaf traits such as leaf dry matter content were response traits strongly influenced by land use and abiotic environment, with follow-on effects on several ecosystem properties (e.g., green biomass production), and could therefore be used as functional markers of ES." AUTHOR'S DESCRIPTION: "Variation in green biomass production was modelled using…traits community-weighted mean (CWM) and functional divergence (FD) and abiotic variables (continuous variables; trait + abiotic) following Diaz et al. (2007). …The comparison between this model and the land-use alone model identifies the need for site-based information beyond a land use or land cover proxy, and the comparison with the land use + abiotic model assesses the value of additional ecological (trait) information…Green biomass production for each pixel was calculated and mapped using model estimates for…regression coefficients on abiotic variables and traits. For each pixel these calculations were applied to mapped estimates of abiotic variables and trait CWM and FD. This step is critically novel as compared to a direct application of the model by Diaz et al. (2007) in that we explicitly modelled the responses of trait community-weighted means and functional divergences to environment prior to evaluating their effects on ecosystem properties. Such an approach is the key to the explicit representation of functional variation across the landscape, as opposed to the use of unique trait values within each land use (see Albert et al. 2010)." | ABSTRACT: "Here, we propose a new approach for the analysis, mapping and understanding of multiple ES delivery in landscapes. Spatially explicit single ES models based on plant traits and abiotic characteristics are combined to identify ‘hot’ and ‘cold’ spots of multiple ES delivery, and the land use and biotic determinants of such distributions. We demonstrate the value of this trait-based approach as compared to a pure land-use approach for a pastoral landscape from the central French Alps, and highlight how it improves understanding of ecological constraints to, and opportunities for, the delivery of multiple services." AUTHOR'S DESCRIPTION: "Community-weighted mean date of flowering onset was modelled using mixed models with land use and abiotic variables as fixed effects (LU + abiotic model) and year as a random effect…and modelled for each 20 x 20 m pixel using GLM estimated effects for each land use category and estimated regression coefficients with abiotic variables." | ABSTRACT: "Here, we propose a new approach for the analysis, mapping and understanding of multiple ES delivery in landscapes. Spatially explicit single ES models based on plant traits and abiotic characteristics are combined to identify ‘hot’ and ‘cold’ spots of multiple ES delivery, and the land use and biotic determinants of such distributions. We demonstrate the value of this trait-based approach as compared to a pure land-use approach for a pastoral landscape from the central French Alps, and highlight how it improves understanding of ecological constraints to, and opportunities for, the delivery of multiple services. Vegetative height and leaf traits such as leaf dry matter content were response traits strongly influenced by land use and abiotic environment, with follow-on effects on several ecosystem properties, and could therefore be used as functional markers of ES." AUTHOR'S DESCRIPTION: "Functional divergence of flowering date was modelled using mixed models with land use and abiotic variables as fixed effects (LU + abiotic model) and year as a random effect…and modelled for each 20 x 20 m pixel using GLM estimated effects for each land use category and estimated regression coefficients with abiotic variables." | ABSTRACT: "A price on carbon is expected to generate demand for carbon offset schemes. This demand could drive investment in tree-based monocultures that provide higher carbon yields than diverse plantings of native tree and shrub species, which sequester less carbon but provide greater variation in vegetation structure and composition. Economic instruments such as species conservation banking, the creation and trading of credits that represent biological-diversity values on private land, could close the financial gap between monocultures and more diverse plantings by providing payments to individuals who plant diverse species in locations that contribute to conservation and restoration goals. We studied a highly modified agricultural system in southern Australia that is typical of many temperate agriculture zones globally (i.e., has a high proportion of endangered species, high levels of habitat fragmentation, and presence of non-native species). We quantified the economic returns...from carbon plantings (monoculture and mixed tree and shrubs) under six carbon-price scenarios." AUTHOR'S DESCRIPTION: "The economic returns of carbon plantings are highly variable and depend primarily on carbon yield and price and opportunity costs (Newell & Stavins 2000; Richards & Stokes 2004; Torres et al. 2010)...The spatial variation in carbon yield and costs, including establishment, maintenance, transaction, and opportunity costs, means that the net economic returns of carbon plantings are also likely to vary spatially." | ABSTRACT: "We develop a novel, mechanistic landscape model for biological control of cereal aphids, explicitly accounting for the influence of landscape composition on natural enemies varying in mobility, feeding rates and other life history traits. Finally, we use the model to map biological control services across cereal fields in a Swedish agricultural region with varying landscape complexity. The model predicted that biological control would reduce crop damage by 45–70% and that the biological control effect would be higher in complex landscapes. In a validation with independent data, the model performed well and predicted a significant proportion of biological control variation in cereal fields. However, much variability remains to be explained, and we propose that the model could be improved by refining the mechanistic understanding of predator dynamics and accounting for variation in aphid colonization." |
Specific Policy or Decision Context Cited
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None identified | None identified | None identified | None identified | None identified |
Biophysical Context
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Elevation ranges from 1552 to 2442 m, on predominately south-facing slopes | Elevation ranges from 1552 to 2442 m, on predominantly south-facing slopes | Elevations ranging from 1552 m to 2442 m, on predominantly south-facing slopes | Mix of remnant native vegetation and agricultural land. Remnant vegetation is in 20 large (>10,000 ha) contiguous fragments where rainfall is low. Acacia spp. and Eucalyptus spp. are the dominant tree species in the remnant vegetation, and major native vegetation types are open forests, woodlands, and open woodlands. Dominant agricultural uses are annual crops, annual legumes, and grazing of sheep and cows. The climate is Mediterranean with average annual rainfall ranging from 250 mm to 1000 mm. | Spring-sown cereal croplands, where the bird chearry-oat aphid is a key aphid pest. The aphid colonizes the crop during late May and early June, depending on weather and location. The colonization phase is followed by a brief phase of rapid exponential population growth by wingless aphids, continuing until about the time of crop heading, in late June or early July. After heading, aphid populations usually decline rapidly in the crop due to decreased plant quality and migration to grasslands. The aphids are attacked by a complex of arthropod natural enemies, but parasitism is not important in the region and therefore not modelled here. |
EM Scenario Drivers
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No scenarios presented | No scenarios presented | No scenarios presented | Carbon prices at $10/t CO2^-e, $15/t CO2^-e, $20/t CO2^-e, $25/t CO2^-e, $30/t CO2^-e, and $40/t CO2^-e | No scenarios presented |
EM ID
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EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
Method Only, Application of Method or Model Run
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Method + Application | Method + Application | Method + Application |
Method + Application (multiple runs exist) View EM Runs ?Comment:Runs are differentiated based on the the expected annual profit from two types of carbon plantings: 1) Tree-based monocultures (i.e., monoculture carbon planting) and 2) Diverse plantings of native tree and shrub species (i.e., ecological carbon planting) |
Method + Application |
New or Pre-existing EM?
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New or revised model | New or revised model | New or revised model | New or revised model | New or revised model |
Related EMs (for example, other versions or derivations of this EM) described in ESML
EM ID
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EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
Document ID for related EM
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Doc-260 | Doc-260 | Doc-269 | Doc-260 | Doc-269 | Doc-245 | Doc-246 | Doc-247 | Doc-243 | None |
EM ID for related EM
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EM-66 | EM-68 | EM-69 | EM-70 | EM-71 | EM-79 | EM-80 | EM-81 | EM-82 | EM-83 | EM-65 | EM-66 | EM-68 | EM-69 | EM-70 | EM-79 | EM-80 | EM-81 | EM-82 | EM-83 | EM-65 | EM-66 | EM-68 | EM-69 | EM-70 | EM-71 | EM-80 | EM-81 | EM-82 | EM-83 | EM-128 | EM-141 | None |
EM Modeling Approach
EM ID
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EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
EM Temporal Extent
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2007-2009 | 2007-2008 | 2007-2008 | 2009-2050 | 2009 |
EM Time Dependence
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time-stationary | time-stationary | time-stationary | time-dependent | time-stationary |
EM Time Reference (Future/Past)
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Not applicable | Not applicable | Not applicable | future time | Not applicable |
EM Time Continuity
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Not applicable | Not applicable | Not applicable | discrete | Not applicable |
EM Temporal Grain Size Value
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Not applicable | Not applicable | Not applicable | 1 | Not applicable |
EM Temporal Grain Size Unit
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Not applicable | Not applicable | Not applicable | Year | Not applicable |
EM ID
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EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
Bounding Type
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Physiographic or Ecological | Physiographic or Ecological | Physiographic or Ecological | Physiographic or Ecological | Geopolitical |
Spatial Extent Name
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Central French Alps | Central French Alps | Central French Alps | Agricultural districts of the state of South Australia | Uppland province |
Spatial Extent Area (Magnitude)
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10-100 km^2 | 10-100 km^2 | 10-100 km^2 | 100,000-1,000,000 km^2 | 10,000-100,000 km^2 |
EM ID
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EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
EM Spatial Distribution
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spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially distributed (in at least some cases) |
Spatial Grain Type
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area, for pixel or radial feature | area, for pixel or radial feature | area, for pixel or radial feature | area, for pixel or radial feature | area, for pixel or radial feature |
Spatial Grain Size
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20 m x 20 m | 20 m x 20 m | 20 m x 20 m | 1 ha x 1 ha | 25 m x 25 m |
EM ID
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EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
EM Computational Approach
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Analytic | Analytic | Analytic | Analytic | Analytic |
EM Determinism
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deterministic | deterministic | deterministic | deterministic | deterministic |
Statistical Estimation of EM
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EM ID
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EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
Model Calibration Reported?
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No | No | No | No | No |
Model Goodness of Fit Reported?
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Yes | Yes | Yes | No | No |
Goodness of Fit (metric| value | unit)
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None | None |
Model Operational Validation Reported?
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Yes | No | No | No | Yes |
Model Uncertainty Analysis Reported?
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No | No | No | No | No |
Model Sensitivity Analysis Reported?
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No | No | No | No |
Yes ?Comment:AUTHOR'S NOTE: "Varying aphid fecundity, overall predator abundances and attack rates affected the biological control effect, but had little influence on the relative differences between landscapes with high and low levels of biological control. The model predictions were more sensitive to changing the predators' landscape relations, but, with few exceptions, did not dramatically alter the overall patterns generated by the model." |
Model Sensitivity Analysis Include Interactions?
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Not applicable | Not applicable | Not applicable | Not applicable | No |
EM Locations, Environments, Ecology
Terrestrial location (Classification hierarchy: Continent > Country > U.S. State [United States only])
EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
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Marine location (Classification hierarchy: Realm > Region > Province > Ecoregion)
EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
None | None | None | None | None |
Centroid Lat/Long (Decimal Degree)
EM ID
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EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
Centroid Latitude
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45.05 | 45.05 | 45.05 | -34.9 | 59.52 |
Centroid Longitude
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6.4 | 6.4 | 6.4 | 138.7 | 17.9 |
Centroid Datum
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WGS84 | WGS84 | WGS84 | WGS84 | WGS84 |
Centroid Coordinates Status
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Provided | Provided | Provided | Estimated | Estimated |
EM ID
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EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
EM Environmental Sub-Class
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Agroecosystems | Grasslands | Agroecosystems | Grasslands | Agroecosystems | Grasslands | Agroecosystems | Agroecosystems | Grasslands |
Specific Environment Type
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Subalpine terraces, grasslands, and meadows | Subalpine terraces, grasslands, and meadows. | Subalpine terraces, grasslands, and meadows | Agricultural land for annual crops, annual legumes, and grazing of sheep and cows | Spring-sown cereal croplands and surrounding grassland and non-arable land |
EM Ecological Scale
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Not applicable | Not applicable | Ecological scale is coarser than that of the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class |
Scale of differentiation of organisms modeled
EM ID
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EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
EM Organismal Scale
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Community | Community | Community | Guild or Assemblage | Individual or population, within a species |
Taxonomic level and name of organisms or groups identified
EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
None Available | None Available | None Available |
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EnviroAtlas URL
EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
GAP Ecological Systems | None Available | None Available | Carbon Storage by Tree Biomass | GAP Ecological Systems |
EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
CICES v 4.3 - Common International Classification of Ecosystem Services (Section > Division > Group > Class)
EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
None | None | None |
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<a target="_blank" rel="noopener noreferrer" href="https://www.epa.gov/eco-research/national-ecosystem-services-classification-system-nescs-plus">National Ecosystem Services Classification System (NESCS) Plus</a>
(Environmental Subclass > Ecological End-Product (EEP) > EEP Subclass > EEP Modifier)
EM-65 | EM-71 | EM-79 |
EM-127 ![]() |
EM-303 |
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None | None | None |
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