EcoService Models Library (ESML)
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Compare EMs
Which comparison is best for me?EM Variables by Variable Role
One quick way to compare ecological models (EMs) is by comparing their variables. Predictor variables show what kinds of influences a model is able to account for, and what kinds of data it requires. Response variables show what information a model is capable of estimating.
This first comparison shows the names (and units) of each EM’s variables, side-by-side, sorted by variable role. Variable roles in ESML are as follows:
- Predictor Variables
- Time- or Space-Varying Variables
- Constants and Parameters
- Intermediate (Computed) Variables
- Response Variables
- Computed Response Variables
- Measured Response Variables
EM Variables by Category
A second way to use variables to compare EMs is by focusing on the kind of information each variable represents. The top-level categories in the ESML Variable Classification Hierarchy are as follows:
- Policy Regarding Use or Management of Ecosystem Resources
- Land Surface (or Water Body Bed) Cover, Use or Substrate
- Human Demographic Data
- Human-Produced Stressor or Enhancer of Ecosystem Goods and Services Production
- Ecosystem Attributes and Potential Supply of Ecosystem Goods and Services
- Non-monetary Indicators of Human Demand, Use or Benefit of Ecosystem Goods and Services
- Monetary Values
Besides understanding model similarities, sorting the variables for each EM by these 7 categories makes it easier to see if the compared models can be linked using similar variables. For example, if one model estimates an ecosystem attribute (in Category 5), such as water clarity, as a response variable, and a second model uses a similar attribute (also in Category 5) as a predictor of recreational use, the two models can potentially be used in tandem. This comparison makes it easier to spot potential model linkages.
All EM Descriptors
This selection allows a more detailed comparison of EMs by model characteristics other than their variables. The 50-or-so EM descriptors for each model are presented, side-by-side, in the following categories:
- EM Identity and Description
- EM Modeling Approach
- EM Locations, Environments, Ecology
- EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
EM Descriptors by Modeling Concepts
This feature guides the user through the use of the following seven concepts for comparing and selecting EMs:
- Conceptual Model
- Modeling Objective
- Modeling Context
- Potential for Model Linkage
- Feasibility of Model Use
- Model Certainty
- Model Structural Information
Though presented separately, these concepts are interdependent, and information presented under one concept may have relevance to other concepts as well.
EM Identity and Description
EM ID
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EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
EM Short Name
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Community flowering date, Central French Alps | Coral taxa and land development, St.Croix, VI, USA | Coastal protection in Belize | Seed mix for native plant establishment, IA, USA | HWB Blood pressure, Great Lakes waterfront, USA | ESTIMAP - Pollination potential, Iran | EcoSim II - method |
EM Full Name
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Community weighted mean flowering date, Central French Alps | Coral taxa richness and land development, St.Croix, Virgin Islands, USA | Coastal Protection provided by Coral, Seagrasses and Mangroves in Belize: | Cost-effective seed mix design for native plant establishment, Iowa, USA | Human well being indicator- Blood pressure, Great Lakes waterfront, USA | ESTIMAP - Pollination potential, Iran | EcoSim II - method |
EM Source or Collection
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EU Biodiversity Action 5 | US EPA | InVEST | None | None | None | None |
EM Source Document ID
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260 | 96 | 350 | 394 |
422 ?Comment:Has not been submitted to Journal yet, but has been peer reviewed by EPA inhouse and outside reviewers |
434 | 448 |
Document Author
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Lavorel, S., Grigulis, K., Lamarque, P., Colace, M-P, Garden, D., Girel, J., Pellet, G., and Douzet, R. | Oliver, L. M., Lehrter, J. C. and Fisher, W. S. | Guannel, G., Arkema, K., Ruggiero, P., and G. Verutes | Meissen, J. | Ted R. Angradi, Jonathon J. Launspach, and Molly J. Wick | Rahimi, E., Barghjelveh, S., and P. Dong | Walters, C., Pauly, D., Christensen, V., and J.F. Kitchell |
Document Year
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2011 | 2011 | 2016 | 2018 | None | 2020 | 2000 |
Document Title
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Using plant functional traits to understand the landscape distribution of multiple ecosystem services | Relating landscape development intensity to coral reef condition in the watersheds of St. Croix, US Virgin Islands | The Power of Three: Coral Reefs, Seagrasses and Mangroves Protect Coastal Regions and Increase Their Resilience | Cost-effective seed mix design and first-year management | Human well-being and natural capital indictors for Great Lakes waterfront revitalization | Using the Lonsdorf and ESTIMAP models for large-scale pollination Using the Lonsdorf and ESTIMAP models for large-scale pollination mapping (Case study: Iran) | Representing density dependent consequences of life history strategies in aquatic ecostems: EcoSim II |
Document Status
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Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed but unpublished (explain in Comment) | Peer reviewed and published | Peer reviewed and published |
Comments on Status
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Published journal manuscript | Published journal manuscript | Published journal manuscript | Published report | Journal manuscript submitted or in review | Published journal manuscript | Published journal manuscript |
EM ID
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EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
Not applicable | Not applicable | Not identified in paper | Not applicable | Not applicable | Not applicable | https://ecopath.org/downloads/ | |
Contact Name
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Sandra Lavorel | Leah Oliver | Greg Guannel | Justin Meissen | Ted Angradi | Ehsan Rahini | Carl Walters |
Contact Address
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Laboratoire d’Ecologie Alpine, UMR 5553 CNRS Université Joseph Fourier, BP 53, 38041 Grenoble Cedex 9, France | National Health and Environmental Research Effects Laboratory | The Nature Conservancy, Coral Gables, FL. USA | Tallgrass Prairie Center, University of Northern Iowa | USEPA, Center for Computational Toxicology and Ecology, Great Lakes Toxicology and Ecology Division, Duluth, MN 55804 | Environmental Sciences Research Institute, Shahid Beheshti University, Tehran, Iran | Fisheries Centre, University of British Columbia, Vancouver, British Columbia, British Columbia, Canada, V6T 1Z4 |
Contact Email
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sandra.lavorel@ujf-grenoble.fr | leah.oliver@epa.gov | greg.guannel@gmail.com | Not reported | tedangradi@gmail.com | ehsanrahimi666@gmail.com | c.walters@oceans.ubc.ca |
EM ID
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EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
Summary Description
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ABSTRACT: "Here, we propose a new approach for the analysis, mapping and understanding of multiple ES delivery in landscapes. Spatially explicit single ES models based on plant traits and abiotic characteristics are combined to identify ‘hot’ and ‘cold’ spots of multiple ES delivery, and the land use and biotic determinants of such distributions. We demonstrate the value of this trait-based approach as compared to a pure land-use approach for a pastoral landscape from the central French Alps, and highlight how it improves understanding of ecological constraints to, and opportunities for, the delivery of multiple services." AUTHOR'S DESCRIPTION: "Community-weighted mean date of flowering onset was modelled using mixed models with land use and abiotic variables as fixed effects (LU + abiotic model) and year as a random effect…and modelled for each 20 x 20 m pixel using GLM estimated effects for each land use category and estimated regression coefficients with abiotic variables." | AUTHOR'S DESCRIPTION: "In this exploratory comparison, stony coral condition was related to watershed LULC and LDI values. We also compared the capacity of other potential human activity indicators to predict coral reef condition using multivariate analysis." (294) | AUTHOR'S DESCRIPTION: "Natural habitats have the ability to protect coastal communities against the impacts of waves and storms, yet it is unclear how different habitats complement each other to reduce those impacts. Here, we investigate the individual and combined coastal protection services supplied by live corals on reefs, seagrass meadows, and mangrove forests during both non-storm and storm conditions, and under present and future sea-level conditions. Using idealized profiles of fringing and barrier reefs, we quantify the services supplied by these habitats using various metrics of inundation and erosion. We find that, together, live corals, seagrasses, and mangroves supply more protection services than any individual habitat or any combination of two habitats. Specifically, we find that, while mangroves are the most effective at protecting the coast under non-storm and storm conditions, live corals and seagrasses also moderate the impact of waves and storms, thereby further reducing the vulnerability of coastal regions. Also, in addition to structural differences, the amount of service supplied by habitats in our analysis is highly dependent on the geomorphic setting, habitat location and forcing conditions: live corals in the fringing reef profile supply more protection services than seagrasses; seagrasses in the barrier reef profile supply more protection services than live corals; and seagrasses, in our simulations, can even compensate for the long-term degradation of the barrier reef. Results of this study demonstrate the importance of taking integrated and place-based approaches when quantifying and managing for the coastal protection services supplied by ecosystems." | AUTHOR'S DESCRIPTION: "Restoring ecosystem services at scale requires executing conservation programs in a way that is resource and cost efficient as well as ecologically effective…Seed mix design is one of the largest determinants of project cost and ecological outcomes for prairie reconstructions. In particular, grass-to-forb seeding ratio affects cost since forb seed can be much more expensive relative to grass species (Prairie Moon Nursery 2012). Even for seed mixes with the same overall seeding rates, a mix with a low grass-to-forb seeding ratio is considerably more expensive than one with a high grass-to-forb ratio. Seeding rates for different plant functional groups that are too high or low may also adversely affect ecological outcomes…First-year management may also play a role in cost-effective prairie reconstruction. Post-agricultural sites where restoration typically occurs are often quickly dominated by fast-growing annual weeds by the time sown prairie seeds begin germinating (Smith et al. 2010)… Williams and others (2007) showed that prairie seedlings sown into established warm-season grasses were reliant on high light conditions created by frequently mowing tall vegetation in order to survive in subsequent years…Our objective was to compare native plant establishment and cost effectiveness with and without first-year mowing for three different seed mixes that differed in grass to forb ratio and soil type customization. With knowledge of plant establishment, cost effectiveness, and mowing management outcomes, conservation practitioners will be better equipped to restore prairie efficiently and successfully." | ABSTRACT: "Revitalization of natural capital amenities at the Great Lakes waterfront can result from sediment remediation, habitat restoration, climate resilience projects, brownfield reuse, economic redevelopment and other efforts. Practical indicators are needed to assess the socioeconomic and cultural benefits of these investments. We compiled U.S. census-tract scale data for five Great Lakes communities: Duluth/Superior, Green Bay, Milwaukee, Chicago, and Cleveland. We downloaded data from the US Census Bureau, Centers for Disease Control and Prevention, Environmental Protection Agency, National Oceanic and Atmospheric Administration, and non-governmental organizations. We compiled a final set of 19 objective human well-being (HWB) metrics and 26 metrics representing attributes of natural and 7 seminatural amenities (natural capital). We rated the reliability of metrics according to their consistency of correlations with metric of the other type (HWB vs. natural capital) at the census-tract scale, how often they were correlated in the expected direction, strength of correlations, and other attributes. Among the highest rated HWB indicators were measures of mean health, mental health, home ownership, home value, life success, and educational attainment. Highest rated natural capital metrics included tree cover and impervious surface metrics, walkability, density of recreational amenities, and shoreline type. Two ociodemographic covariates, household income and population density, had a strong influence on the associations between HWB and natural capital and must be included in any assessment of change in HWB benefits in the waterfront setting. Our findings are a starting point for applying objective HWB and natural capital indicators in a waterfront revitalization context." | Abstract: ". ..we used the ESTIMAP model to improve the results of the Lonsdorf model. For this, we included the effects of roads, railways, rivers, wetlands, lakes, altitude, climate, and ecosystem boundaries in the ESTIMAP modeling and compared the results with the Lonsdorf model. The results of the Lonsdorf model showed that the majority of Iran had a very low potential for providing pollination service and only three percent of the northern and western parts of Iran had high potential. However, the results of the ESTIMAP model showed that 16% of Iran had a high potential to provide pollination that covers most of the northern and southern parts of the country. The results of the ESTIMAP model for pollination mapping in Iran showed the Lonsdorf model of estimating pollination service can be improved through considering other relevant factors." | ABSTRACT: " EcoSim II uses results from the Ecopath procedure for trophic mass-balance analysis to define biomass dynamics models for predicting temporal change in exploited ecosystems. Key populations can be repre- sented in further detail by using delay-difference models to account for both biomass and numbers dynamics. A major problem revealed by linking the population and biomass dynamics models is in representation of population responses to changes in food supply; simple proportional growth and reproductive responses lead to unrealistic predic- tions of changes in mean body size with changes in fishing mortality. EcoSim II allows users to specify life history mechanisms to avoid such unrealistic predictions: animals may translate changes in feed- ing rate into changes in reproductive rather than growth rates, or they may translate changes in food availability into changes in foraging time that in turn affects predation risk. These options, along with model relationships for limits on prey availabil- ity caused by predation avoidance tactics, tend to cause strong compensatory responses in modeled populations. It is likely that such compensatory responses are responsible for our inability to find obvious correlations between interacting trophic components in fisheries time-series data. But Eco- sim II does not just predict strong compensatory responses: it also suggests that large piscivores may be vulnerable to delayed recruitment collapses caused by increases in prey species that are in turn competitors/predators of juvenile piscivores " |
Specific Policy or Decision Context Cited
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None identified | Not applicable | Future rock lobster fisheries management | Seed mix design and management practices for native plant restoration | None identified | None reported | None |
Biophysical Context
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Elevation ranges from 1552 to 2442 m, on predominantly south-facing slopes | nearshore; <1.5 km offshore; <12 m depth | barrier reef and fringing reef in nearshore coastal marine system | The soils underlying the study site are primarily poorly drained Clyde clay loams, with a minor component of somewhat poorly drained Floyd loams in the northwest (NRCS 2016). Topographically, the study site is level, and slopes do not exceed 5% grade. Land use prior to this experiment was agricultural, with corn and soybeans consistently grown in rotation at the site. | Waterfront districts on south Lake Michigan and south lake Erie | None additional | None, Ocean ecosystems |
EM Scenario Drivers
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No scenarios presented | Not applicable | Reef type, Sea level increase, storm conditions, seagrass conditions, coral conditions, vegetation types and conditions | No scenarios presented | N/A | N/A | N/A |
EM ID
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EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
Method Only, Application of Method or Model Run
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Method + Application | Method + Application | Method + Application (multiple runs exist) View EM Runs | Method + Application (multiple runs exist) View EM Runs | Method + Application | Method + Application | Method Only |
New or Pre-existing EM?
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New or revised model | New or revised model | New or revised model | New or revised model | New or revised model | Application of existing model | New or revised model |
Related EMs (for example, other versions or derivations of this EM) described in ESML
EM ID
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EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
Document ID for related EM
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Doc-260 | Doc-269 | None | None | Doc-395 | Doc-422 | Doc-432 | None |
EM ID for related EM
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EM-65 | EM-66 | EM-68 | EM-69 | EM-70 | EM-79 | EM-80 | EM-81 | EM-82 | EM-83 | None | None | EM-728 | EM-886 | EM-888 | EM-889 | EM-891 | EM-893 | EM-894 | EM-895 | EM-939 | None |
EM Modeling Approach
EM ID
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EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
EM Temporal Extent
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2007-2008 | 2006-2007 | 2005-2013 | 2015-2017 | 2022 | 2020 | Not applicable |
EM Time Dependence
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time-stationary | time-stationary | time-dependent | time-dependent | time-stationary | time-stationary | time-dependent |
EM Time Reference (Future/Past)
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Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | both |
EM Time Continuity
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Not applicable | Not applicable | discrete | discrete | Not applicable | Not applicable |
discrete ?Comment:Modeller dependent |
EM Temporal Grain Size Value
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Not applicable | Not applicable | 1 | 1 | Not applicable | Not applicable | 1 |
EM Temporal Grain Size Unit
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Not applicable | Not applicable | Second | Year | Not applicable | Not applicable | Day |
EM ID
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EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
Bounding Type
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Physiographic or Ecological | Physiographic or Ecological | Geopolitical | Other | Geopolitical | Geopolitical | Other |
Spatial Extent Name
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Central French Alps | St.Croix, U.S. Virgin Islands | Coast of Belize | Iowa State University Northeast Research and Demonstration Farm | Great Lakes waterfront | Iran | Not applicable |
Spatial Extent Area (Magnitude)
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10-100 km^2 | 10-100 km^2 | 100-1000 km^2 | <1 ha | 1000-10,000 km^2. | >1,000,000 km^2 | Not applicable |
EM ID
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EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
EM Spatial Distribution
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spatially distributed (in at least some cases) | spatially lumped (in all cases) | spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially lumped (in all cases) |
spatially distributed (in at least some cases) ?Comment:Varies by inputs, but results are for areas of country |
spatially lumped (in all cases) |
Spatial Grain Type
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area, for pixel or radial feature | Not applicable | length, for linear feature (e.g., stream mile) | area, for pixel or radial feature | Not applicable | area, for pixel or radial feature | Not applicable |
Spatial Grain Size
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20 m x 20 m | Not applicable | 1 meter | 20 ft x 28 ft | Not applicable | ha^2 | Not applicable |
EM ID
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EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
EM Computational Approach
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Analytic | Analytic | Analytic | Analytic | Numeric | Numeric | Analytic |
EM Determinism
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deterministic | deterministic | deterministic | stochastic | deterministic | deterministic | deterministic |
Statistical Estimation of EM
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EM ID
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EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
Model Calibration Reported?
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No | Yes | No | Not applicable | No | No | No |
Model Goodness of Fit Reported?
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Yes | Yes | No | Not applicable | No | No | No |
Goodness of Fit (metric| value | unit)
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None | None | None | None | None |
Model Operational Validation Reported?
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No | No |
No ?Comment:Used the SWAN model (see below for referenece) with Generation 1 or 2 wind-wave formulations to validate the wave development portion of the model. Booij N, Ris RC, Holthuijsen LH. A third-generation wave model for coastal regions 1. Model description and validation. J Geophys Res. American Geophysical Union; 1999;104: 7649?7666. |
No | No | No | Not applicable |
Model Uncertainty Analysis Reported?
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No | Yes | No | Not applicable | No | No | Not applicable |
Model Sensitivity Analysis Reported?
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No | No | No | Not applicable | Yes | No | Not applicable |
Model Sensitivity Analysis Include Interactions?
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Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable |
EM Locations, Environments, Ecology
Terrestrial location (Classification hierarchy: Continent > Country > U.S. State [United States only])
EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
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None |
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Comment:Model for Iran - no form preset id for country |
None |
Marine location (Classification hierarchy: Realm > Region > Province > Ecoregion)
EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
None |
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None | None | None | None |
Centroid Lat/Long (Decimal Degree)
EM ID
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EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
Centroid Latitude
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45.05 | 17.75 | 18.63 | 42.93 | 42.26 | 32.29 | Not applicable |
Centroid Longitude
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6.4 | -64.75 | -88.22 | -92.57 | -87.84 | 53.68 | Not applicable |
Centroid Datum
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WGS84 | NAD83 | WGS84 | WGS84 | WGS84 | WGS84 | Not applicable |
Centroid Coordinates Status
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Provided | Estimated | Estimated | Provided | Estimated | Estimated | Not applicable |
EM ID
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EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
EM Environmental Sub-Class
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Agroecosystems | Grasslands | Near Coastal Marine and Estuarine | Near Coastal Marine and Estuarine | Agroecosystems | Grasslands | Terrestrial Environment (sub-classes not fully specified) | Terrestrial Environment (sub-classes not fully specified) | Open Ocean and Seas |
Specific Environment Type
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Subalpine terraces, grasslands, and meadows. | stony coral reef | coral reefs | Research farm in historic grassland | Lake Michigan & Lake Erie waterfront | terrestrial land types | Pelagic |
EM Ecological Scale
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Not applicable | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class |
Scale of differentiation of organisms modeled
EM ID
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EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
EM Organismal Scale
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Community | Guild or Assemblage | Guild or Assemblage | Community | Not applicable | Not applicable |
Other (Comment) ?Comment:Varied levels of taxonomic order |
Taxonomic level and name of organisms or groups identified
EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
None Available |
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None Available | None Available | None Available |
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EnviroAtlas URL
EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
CICES v 4.3 - Common International Classification of Ecosystem Services (Section > Division > Group > Class)
EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
None |
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None |
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<a target="_blank" rel="noopener noreferrer" href="https://www.epa.gov/eco-research/national-ecosystem-services-classification-system-nescs-plus">National Ecosystem Services Classification System (NESCS) Plus</a>
(Environmental Subclass > Ecological End-Product (EEP) > EEP Subclass > EEP Modifier)
EM-71 | EM-260 |
EM-542 ![]() |
EM-719 ![]() |
EM-890 | EM-941 | EM-964 |
None |
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None |
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