EcoService Models Library (ESML)
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Compare EMs
Which comparison is best for me?EM Variables by Variable Role
One quick way to compare ecological models (EMs) is by comparing their variables. Predictor variables show what kinds of influences a model is able to account for, and what kinds of data it requires. Response variables show what information a model is capable of estimating.
This first comparison shows the names (and units) of each EM’s variables, side-by-side, sorted by variable role. Variable roles in ESML are as follows:
- Predictor Variables
- Time- or Space-Varying Variables
- Constants and Parameters
- Intermediate (Computed) Variables
- Response Variables
- Computed Response Variables
- Measured Response Variables
EM Variables by Category
A second way to use variables to compare EMs is by focusing on the kind of information each variable represents. The top-level categories in the ESML Variable Classification Hierarchy are as follows:
- Policy Regarding Use or Management of Ecosystem Resources
- Land Surface (or Water Body Bed) Cover, Use or Substrate
- Human Demographic Data
- Human-Produced Stressor or Enhancer of Ecosystem Goods and Services Production
- Ecosystem Attributes and Potential Supply of Ecosystem Goods and Services
- Non-monetary Indicators of Human Demand, Use or Benefit of Ecosystem Goods and Services
- Monetary Values
Besides understanding model similarities, sorting the variables for each EM by these 7 categories makes it easier to see if the compared models can be linked using similar variables. For example, if one model estimates an ecosystem attribute (in Category 5), such as water clarity, as a response variable, and a second model uses a similar attribute (also in Category 5) as a predictor of recreational use, the two models can potentially be used in tandem. This comparison makes it easier to spot potential model linkages.
All EM Descriptors
This selection allows a more detailed comparison of EMs by model characteristics other than their variables. The 50-or-so EM descriptors for each model are presented, side-by-side, in the following categories:
- EM Identity and Description
- EM Modeling Approach
- EM Locations, Environments, Ecology
- EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
EM Descriptors by Modeling Concepts
This feature guides the user through the use of the following seven concepts for comparing and selecting EMs:
- Conceptual Model
- Modeling Objective
- Modeling Context
- Potential for Model Linkage
- Feasibility of Model Use
- Model Certainty
- Model Structural Information
Though presented separately, these concepts are interdependent, and information presented under one concept may have relevance to other concepts as well.
EM Identity and Description
EM ID
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EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
EM Short Name
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InVEST - Water Yield (v3.0) | Reef dive site favorability, St. Croix, USVI | InVEST fisheries, lobster, South Africa | Plant-pollinator networks at reclaimed mine, USA | VELMA v. 2.0 disturbance |
EM Full Name
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InVEST v3.0 Reservoir Hydropower Projection, aka Water Yield | Dive site favorability (reef), St. Croix, USVI | Integrated Valuation of Ecosystem Services and Trade-offs Fisheries, rock lobster, South Africa | Restoration of plant-pollinator networks at reclaimed strip mine, Ohio, USA | VELMA (Visualizing Ecosystems for Land Management Assessment) version 2.0 disturbance |
EM Source or Collection
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InVEST | US EPA | InVEST | None | US EPA |
EM Source Document ID
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311 | 335 |
349 ?Comment:Supplemented with the InVEST Users Guide fisheries. |
397 | 366 |
Document Author
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Natural Capital Project | Yee, S. H., Dittmar, J. A., and L. M. Oliver | Ward, Michelle, Hugh Possingham, Johathan R. Rhodes, Peter Mumby | Cusser, S. and K. Goodell | McKane, R. B., A. Brookes, K. Djang, M. Stieglitz, A. G. Abdelnour, F. Pan, J. J. Halama, P. B. Pettus and D. L. Phillips |
Document Year
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2015 | 2014 | 2018 | 2013 | 2014 |
Document Title
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Water Yield: Reservoir Hydropower Production- InVEST (v3.0) | Comparison of methods for quantifying reef ecosystem services: A case study mapping services for St. Croix, USVI | Food, money and lobsters: Valuing ecosystem services to align environmental management with Sustainable Development Goals | Diversity and distribution of floral resources influence the restoration of plant-pollinator networks on a reclaimed strip mine | VELMA Version 2.0 User Manual and Technical Documentation |
Document Status
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Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published |
Comments on Status
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Web published | Published journal manuscript | Published journal manuscript | Published journal manuscript | Published report |
EM ID
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EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
https://www.naturalcapitalproject.org/invest/ | Not applicable | https://www.naturalcapitalproject.org/invest/ | Not applicable | https://www.epa.gov/water-research/visualizing-ecosystem-land-management-assessments-velma-model-20 | |
Contact Name
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Natural Capital Project | Susan H. Yee | Michelle Ward |
Sarah Cusser ?Comment:Department of Evolution, Ecology, and Organismal Biology, Ohio State University, 318 West 12th Avenue, Columbus, OH 43202, U.S.A. |
Robert B. McKane |
Contact Address
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371 Serra Mall, Stanford University, Stanford, Ca 94305 | US EPA, Office of Research and Development, NHEERL, Gulf Ecology Division, Gulf Breeze, FL 32561, USA | ARC Centre of Excellence for Environmental Decisions, The University of Queensland, Brisbane, QLD 4072, Australia | Department of Evolution, Ecology, and Behavior, School of Biological Sciences, The University of Texas at Austin, 100 East 24th Street Stop A6500, Austin, TX 78712-1598, U.S.A. | U.S. EPA, Office of Research and Development, National Health and Environmental Effects Research Laboratory, Western Ecology Division, Corvallis, Oregon 97333 |
Contact Email
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invest@naturalcapitalproject.org | yee.susan@epa.gov | m.ward@uq.edu.au | sarah.cusser@gmail.com | mckane.bob@epa.gov |
EM ID
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EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
Summary Description
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Please note: This ESML entry describes an InVEST model version that was current as of 2015. More recent versions may be available at the InVEST website. AUTHOR'S DESCRIPTION: "The InVEST Reservoir Hydropower model estimates the relative contributions of water from different parts of a landscape, offering insight into how changes in land use patterns affect annual surface water yield and hydropower production. Modeling the connections between landscape changes and hydrologic processes is not simple. Sophisticated models of these connections and associated processes (such as the WEAP model) are resource and data intensive and require substantial expertise. To accommodate more contexts, for which data are readily available, InVEST maps and models the annual average water yield from a landscape used for hydropower production, rather than directly addressing the affect of LULC changes on hydropower failure as this process is closely linked to variation in water inflow on a daily to monthly timescale. Instead, InVEST calculates the relative contribution of each land parcel to annual average hydropower production and the value of this contribution in terms of energy production. The net present value of hydropower production over the life of the reservoir also can be calculated by summing discounted annual revenues. The model runs on a gridded map. It estimates the quantity and value of water used for hydropower production from each subwatershed in the area of interest. It has three components, which run sequentially. First, it determines the amount of water running off each pixel as the precipitation less the fraction of the water that undergoes evapotranspiration. The model does not differentiate between surface, subsurface and baseflow, but assumes that all water yield from a pixel reaches the point of interest via one of these pathways. This model then sums and averages water yield to the subwatershed level. The pixel-scale calculations allow us to represent the heterogeneity of key driving factors in water yield such as soil type, precipitation, vegetation type, etc. However, the theory we are using as the foundation of this set of models was developed at the subwatershed to watershed scale. We are only confident in the interpretation of these models at the subwatershed scale, so all outputs are summed and/or averaged to the subwatershed scale. We do continue to provide pixel-scale representations of some outputs for calibration and model-checking purposes only. These pixel-scale maps are not to be interpreted for understanding of hydrological processes or to inform decision making of any kind. | ABSTRACT: "...We investigated and compared a number of existing methods for quantifying ecological integrity, shoreline protection, recreational opportunities, fisheries production, and the potential for natural products discovery from reefs. Methods were applied to mapping potential ecosystem services production around St. Croix, U.S. Virgin Islands. Overall, we found that a number of different methods produced similar predictions." AUTHOR'S DESCRIPTION: "A number of methods have been developed for linking biophysical attributes of reef condition, such as reef structural complexity, fish biomass, or species richness, to provisioning of ecosystem goods and services (Principe et al., 2012). We investigated the feasibility of using existing methods and data for mapping production of reef ecosystem goods and services. We applied these methods toward mapping potential ecosystem goods and services production in St. Croix, U.S. Virgin Islands (USVI)...For each of the five categories of ecosystem services, we chose a suite of models and indices for estimating potential production based on relative ease of implementation, consisting of well-defined parameters, and likely availability of input data, to maximize potential for transferability to other locations. For each method, we assembled the necessary reef condition and environmental data as spatial data layers for St. Croix (Table1). The coastal zone surrounding St. Croix was divided into 10x10 m grid cells, and production functions were applied to quantify ecosystem services provisioning in each grid cell...A number of recreational activities are associated directly or indirectly with coral reefs including scuba diving, snorkeling, surfing, underwater photography, recreational fishing, wildlife viewing, beach sunbathing and swimming, and beachcombing (Principe et al., 2012)…In lieu of surveys of diver opinion, recreational opportunities can also be estimated by actual field data of coral condition at preferred dive sites. A few studies have directly examined links between coral condition and production of recreational opportunities through field monitoring in an attempt to validate perceptions of recreational quality (Pendleton, 1994; Williams and Polunin, 2002; Leeworthy et al., 2004; Leujakand Ormond, 2007; Uyarraetal., 2009). Uyarraetal. (2009) used surveys to determine reef attributes related to diver perceptions of most and least favorite dive sites. Field data was used to narrow down the suite of potential preferred attributes to those that reflected actual site condition. We combined these attributes to form an index of dive site favorability: Dive site favorability = ΣipiRi where pi is the proportion of respondents indicating each attribute i that affected dive enjoyment positively. Ri is the mean relative magnitude of measured variables used to quantify each descriptive attribute i, including ‘fish abundance’ (pi=0.803), quantified by number of fish schools and fish species richness, and | AUTHOR'S DESCRIPTION: "Here we develop a method for assessing future scenarios of environmental management change that improve coastal ecosystem services and thereby, support the success of the SDGs. We illustrate application of the method using a case study of South Africa’s West Coast Rock Lobster fishery within the Table Mountain National Park (TMNP) Marine Protected Area...We calculated the retrospective and current value of the West Coast Rock Lobster fishery using published and unpublished data from various sources and combined the market worth of landed lobster from recreational fishers, small-scale fisheries (SSF), large-scale fisheries (LSF) and poachers. Then using the InVEST tool, we combined data to build scenarios that describe possible futures for the West Coast Rock Lobster fishery (see Table 1). The first scenario, entitled ‘Business as Usual’ (BAU), takes the current situation and most up-to-date data to model the future if harvest continues at the existing rate. The second scenario is entitled ‘Redirect the Poachers’ (RP), which attempts to model implementation of strict management, whereby poaching is minimised from the Marine Protected Area and other economic and nutritional sources are made available through government initiatives. The third scenario, entitled ‘Large Scale Cutbacks’ (LSC), excludes large-scale fisheries from harvesting West Coast Rock Lobster within the TMNP Marine Protected Area." | ABSTRACT: "Plant–pollinator mutualisms are one of the several functional relationships that must be reinstated to ensure the long-term success of habitat restoration projects. These mutualisms are unlikely to reinstate themselves until all of the resource requirements of pollinators have been met. By meeting these requirements, projects can improve their long-term success. We hypothesized that pollinator assemblage and structure and stability of plant–pollinator networks depend both on aspects of the surrounding landscape and of the restoration effort itself. We predicted that pollinator species diversity and network stability would be negatively associated with distance from remnant habitat, but that local floral diversity might rescue pollinator diversity and network stability in locations distant from the remnant. We created plots of native prairie on a reclaimed strip mine in central Ohio, U.S.A. that ranged in floral diversity and isolation from the remnant habitat. We found that the pollinator diversity declined with distance from the remnant habitat. Furthermore, reduced pollinator diversity in low floral diversity plots far from the remnant habitat was associated with loss of network stability. High floral diversity, however, compensated for losses in pollinator diversity in plots far from the remnant habitat through the attraction of generalist pollinators. Generalist pollinators increased network connectance and plant-niche overlap. Asa result, network robustness of high floral diversity plots was independent of isolation. We conclude that the aspects of the restoration effort itself, such as floral community composition, can be successfully tailored to incorporate the restoration of pollinators and improve success given a particular landscape context." | VELMA – Visualizing Ecosystems for Land Management Assessments - is a spatially distributed, eco-hydrological model that links a land surface hydrology model with a terrestrial biogeochemistry model for simulating the integrated responses of vegetation, soil, and water resources to interacting stressors. For example, VELMA can simulate how changes in climate and land use interact to affect soil water storage, surface and subsurface runoff, vertical drainage, evapotranspiration, vegetation and soil carbon and nitrogen dynamics, and transport of nitrate, ammonium, and dissolved organic carbon and nitrogen to water bodies. VELMA differs from other existing eco-hydrology models in its simplicity, flexibility, and theoretical foundation. The model has a user-friendly Graphics User Interface (GUI) for easy input of model parameter values. In addition, advanced visualization of simulation results can enhance understanding of results and underlying concepts. VELMA’s visualization and interactivity features are packaged in an open-source, open-platform programming environment (Java / Eclipse). The development team for VELMA version 2.0 includes Dr. Bob McKane and coworkers at the U.S. Environmental Protection Agency’s Western Ecology Division, Dr. Marc Stieglitz and coworkers at the Georgia Institute of Technology, and Dr. Feifei Pan at the University of North Texas. AUTHOR'S DESCRIPTION: "Understanding how disturbances such as harvest, fire and fertilization affect ecosystem services has been a major motivation in the development of VELMA. For example, how do disturbances such as forest harvest or the application of agronomic fertilizers affect hydrological and biogeochemical processes controlling water quality and quantity, carbon sequestration, production of greenhouse gases, etc.? Abdelnour et al. (2011, 2013) have already demonstrated the use of VELMA v1.0 to simulate the effects of forest clearcutting on ecohydrological processes that regulate a variety of ecosystem services. With the addition of a tissue-specific plant biomass (LSR) simulator and an enhanced GUI, VELMA v2.0 significantly expands the detail, flexibility, and ease of use for simulating disturbance effects. Currently available disturbance models include: - BurnDisturbanceModel, effects of fire. - GrazeDisturbanceModel, effects of grazing. - FertilizeLsrDisturbanceModel, effects of fertilizer applications. - HarvestLsrDisturbanceModel, effects of biomass harvest. Each of these disturbance models specifies where and when a disturbance event will occur. The Burn, Graze and Harvest models have options for specifying how much of each plant tissue and detritus pool (leaves, stems, roots) will be removed and where it goes (offsite and/or to a specified onsite C and N pools). The Fertilize model has options for applying nitrogen as ammonium, nitrate, urea and/or manure." |
Specific Policy or Decision Context Cited
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None identified | None identified | Future rock lobster fisheries management | None identified | None identified |
Biophysical Context
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None applicable | No additional description provided | No additional description provided | The site was surface mined for coal until the mid-1980s and soon after recontoured and seeded with a low diversity of non-native grasses and forbes. The property is grassland in a state of arrested succession, unable to support tree growth because of shallow, infertile soils. | No additional description provided |
EM Scenario Drivers
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N/A | No scenarios presented | Fisheries exploitation; fishing vulnerability (of age classes) | No scenarios presented | No scenarios presented |
EM ID
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EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
Method Only, Application of Method or Model Run
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Method Only | Method + Application | Method + Application (multiple runs exist) View EM Runs | Method + Application (multiple runs exist) View EM Runs | Method Only |
New or Pre-existing EM?
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New or revised model | Application of existing model | Application of existing model | New or revised model | New or revised model |
Related EMs (for example, other versions or derivations of this EM) described in ESML
EM ID
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EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
Document ID for related EM
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Doc-307 | Doc-280 | Doc-338 | Doc-205 | None | None | None | Doc-13 | Doc-317 | Doc-366 | Doc-359 |
EM ID for related EM
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EM-437 | EM-148 | EM-344 | EM-111 | None | None | None | EM-883 | EM-884 | EM-375 | EM-379 | EM-380 | EM-605 | EM-892 |
EM Modeling Approach
EM ID
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EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
EM Temporal Extent
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Not applicable | 2006-2007, 2010 | 1986-2115 | 2009-2010 | Not applicable |
EM Time Dependence
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time-dependent | time-stationary | time-dependent | time-stationary | time-dependent |
EM Time Reference (Future/Past)
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future time | Not applicable | future time | Not applicable | Not applicable |
EM Time Continuity
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discrete | Not applicable | discrete | Not applicable | discrete |
EM Temporal Grain Size Value
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1 | Not applicable | 1 | Not applicable | 1 |
EM Temporal Grain Size Unit
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Year | Not applicable | Year | Not applicable | Day |
EM ID
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EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
Bounding Type
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Not applicable | Physiographic or ecological | Geopolitical | Physiographic or ecological | Not applicable |
Spatial Extent Name
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Not applicable | Coastal zone surrounding St. Croix | Table Mountain National Park Marine Protected Area | The Wilds | Not applicable |
Spatial Extent Area (Magnitude)
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Not applicable | 100-1000 km^2 | 100-1000 km^2 | 1-10 km^2 | Not applicable |
EM ID
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EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
EM Spatial Distribution
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spatially distributed (in at least some cases) ?Comment:pixel is likely 30m x 30m |
spatially distributed (in at least some cases) | spatially lumped (in all cases) | spatially distributed (in at least some cases) | spatially distributed (in at least some cases) |
Spatial Grain Type
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area, for pixel or radial feature | area, for pixel or radial feature | Not applicable | area, for pixel or radial feature | area, for pixel or radial feature |
Spatial Grain Size
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Not specified | 10 m x 10 m | Not applicable | 10 m radius | user defined |
EM ID
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EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
EM Computational Approach
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Numeric | Analytic | Numeric | Analytic | Numeric |
EM Determinism
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deterministic | deterministic | deterministic | deterministic | deterministic |
Statistical Estimation of EM
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EM ID
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EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
Model Calibration Reported?
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Yes ?Comment:Annual Yield can be calibrated with actual yield based up 10 year average input data though this was an "optional" part of the model. Calibrate with total precipitation and potential evapotranspiration. Before the calibration process is commenced, the modelers suggest performing a sensitivity analysis with the observed runoff data to define the parameters that influence model outputs the most. The calibration can then focus on highly sensitive parameters followed by less sensitive ones. |
Yes | No | Not applicable | Not applicable |
Model Goodness of Fit Reported?
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Not applicable | No | No | Not applicable | Not applicable |
Goodness of Fit (metric| value | unit)
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None | None | None | None | None |
Model Operational Validation Reported?
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No | Yes |
Yes ?Comment:A validation analysis was carried out running the model using data from 1880 to 2001, and then comparing the output for the adult population with the 2001 published data. |
Yes | Not applicable |
Model Uncertainty Analysis Reported?
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No | No | No | Yes | Not applicable |
Model Sensitivity Analysis Reported?
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Not applicable | No | No | No | Not applicable |
Model Sensitivity Analysis Include Interactions?
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Not applicable | Not applicable | Not applicable | Not applicable | Not applicable |
EM Locations, Environments, Ecology
Terrestrial location (Classification hierarchy: Continent > Country > U.S. State [United States only])
EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
None | None | None |
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None |
Marine location (Classification hierarchy: Realm > Region > Province > Ecoregion)
EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
None |
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None | None |
Centroid Lat/Long (Decimal Degree)
EM ID
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EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
Centroid Latitude
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-9999 | 17.73 | -34.18 | 39.82 | Not applicable |
Centroid Longitude
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-9999 | -64.77 | 18.35 | -81.75 | Not applicable |
Centroid Datum
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Not applicable | WGS84 | WGS84 | WGS84 | Not applicable |
Centroid Coordinates Status
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Not applicable | Estimated | Provided | Provided | Not applicable |
EM ID
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EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
EM Environmental Sub-Class
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Rivers and Streams | Near Coastal Marine and Estuarine | Near Coastal Marine and Estuarine | Grasslands | Terrestrial Environment (sub-classes not fully specified) |
Specific Environment Type
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Watershed | Coral reefs | Rocky coast, mixed coast, sandy coast, rocky inshore, sandy inshore, rocky shelf and unconsolidated shelf | Grassland | Terrestrial environment sub-classes |
EM Ecological Scale
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Not applicable | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class |
Scale of differentiation of organisms modeled
EM ID
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EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
EM Organismal Scale
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Not applicable | Guild or Assemblage | Individual or population, within a species | Species | Not applicable |
Taxonomic level and name of organisms or groups identified
EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
None Available | None Available |
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None Available |
EnviroAtlas URL
EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
CICES v 4.3 - Common International Classification of Ecosystem Services (Section > Division > Group > Class)
EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
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<a target="_blank" rel="noopener noreferrer" href="https://www.epa.gov/eco-research/national-ecosystem-services-classification-system-nescs-plus">National Ecosystem Services Classification System (NESCS) Plus</a>
(Environmental Subclass > Ecological End-Product (EEP) > EEP Subclass > EEP Modifier)
EM-368 | EM-456 |
EM-541 ![]() |
EM-774 ![]() |
EM-887 |
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None | None |