EcoService Models Library (ESML)
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Compare EMs
Which comparison is best for me?EM Variables by Variable Role
One quick way to compare ecological models (EMs) is by comparing their variables. Predictor variables show what kinds of influences a model is able to account for, and what kinds of data it requires. Response variables show what information a model is capable of estimating.
This first comparison shows the names (and units) of each EM’s variables, side-by-side, sorted by variable role. Variable roles in ESML are as follows:
- Predictor Variables
- Time- or Space-Varying Variables
- Constants and Parameters
- Intermediate (Computed) Variables
- Response Variables
- Computed Response Variables
- Measured Response Variables
EM Variables by Category
A second way to use variables to compare EMs is by focusing on the kind of information each variable represents. The top-level categories in the ESML Variable Classification Hierarchy are as follows:
- Policy Regarding Use or Management of Ecosystem Resources
- Land Surface (or Water Body Bed) Cover, Use or Substrate
- Human Demographic Data
- Human-Produced Stressor or Enhancer of Ecosystem Goods and Services Production
- Ecosystem Attributes and Potential Supply of Ecosystem Goods and Services
- Non-monetary Indicators of Human Demand, Use or Benefit of Ecosystem Goods and Services
- Monetary Values
Besides understanding model similarities, sorting the variables for each EM by these 7 categories makes it easier to see if the compared models can be linked using similar variables. For example, if one model estimates an ecosystem attribute (in Category 5), such as water clarity, as a response variable, and a second model uses a similar attribute (also in Category 5) as a predictor of recreational use, the two models can potentially be used in tandem. This comparison makes it easier to spot potential model linkages.
All EM Descriptors
This selection allows a more detailed comparison of EMs by model characteristics other than their variables. The 50-or-so EM descriptors for each model are presented, side-by-side, in the following categories:
- EM Identity and Description
- EM Modeling Approach
- EM Locations, Environments, Ecology
- EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
EM Descriptors by Modeling Concepts
This feature guides the user through the use of the following seven concepts for comparing and selecting EMs:
- Conceptual Model
- Modeling Objective
- Modeling Context
- Potential for Model Linkage
- Feasibility of Model Use
- Model Certainty
- Model Structural Information
Though presented separately, these concepts are interdependent, and information presented under one concept may have relevance to other concepts as well.
EM Identity and Description
EM ID
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EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
EM Short Name
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Green biomass production, Central French Alps | Cultural ecosystem services, Bilbao, Spain | Coral taxa and land development, St.Croix, VI, USA | Coastal protection in Belize | ESTIMAP - Pollination potential, Iran | EcoSim II - method | Velma- 6PPD-Q concentrations, Seattle, WA |
EM Full Name
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Green biomass production, Central French Alps | Cultural ecosystem services, Bilbao, Spain | Coral taxa richness and land development, St.Croix, Virgin Islands, USA | Coastal Protection provided by Coral, Seagrasses and Mangroves in Belize: | ESTIMAP - Pollination potential, Iran | EcoSim II - method | VELMA: 6PPD-Quinone stormwater concentrations , Seattle, Washington |
EM Source or Collection
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EU Biodiversity Action 5 |
None ?Comment:EU Mapping Studies |
US EPA | InVEST | None | None | US EPA |
EM Source Document ID
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260 | 191 | 96 | 350 | 434 | 448 | 465 |
Document Author
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Lavorel, S., Grigulis, K., Lamarque, P., Colace, M-P, Garden, D., Girel, J., Pellet, G., and Douzet, R. | Casado-Arzuaga, I., Onaindia, M., Madariaga, I. and Verburg P. H. | Oliver, L. M., Lehrter, J. C. and Fisher, W. S. | Guannel, G., Arkema, K., Ruggiero, P., and G. Verutes | Rahimi, E., Barghjelveh, S., and P. Dong | Walters, C., Pauly, D., Christensen, V., and J.F. Kitchell | Halama JJ, McKane RB, Barnhart BL, Pettus PP, Brookes AF, Adams AK, Gockel CK, Djang KS, Phan V, Chokshi SM, Graham JJ, Tian Z, Peter KT and Kolodziej,EP |
Document Year
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2011 | 2013 | 2011 | 2016 | 2020 | 2000 | 2024 |
Document Title
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Using plant functional traits to understand the landscape distribution of multiple ecosystem services | Mapping recreation and aesthetic value of ecosystems in the Bilbao Metropolitan Greenbelt (northern Spain) to support landscape planning | Relating landscape development intensity to coral reef condition in the watersheds of St. Croix, US Virgin Islands | The Power of Three: Coral Reefs, Seagrasses and Mangroves Protect Coastal Regions and Increase Their Resilience | Using the Lonsdorf and ESTIMAP models for large-scale pollination Using the Lonsdorf and ESTIMAP models for large-scale pollination mapping (Case study: Iran) | Representing density dependent consequences of life history strategies in aquatic ecostems: EcoSim II | Watershed analysis of urban stormwater contaminant 6PPD-Quinone hotspots and stream concentrations using a process-based ecohydrological model |
Document Status
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Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published |
Comments on Status
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Published journal manuscript | Published journal manuscript | Published journal manuscript | Published journal manuscript | Published journal manuscript | Published journal manuscript | Published journal manuscript |
EM ID
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EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
Not applicable | Not applicable | Not applicable | Not identified in paper | Not applicable | https://ecopath.org/downloads/ | Not reported | |
Contact Name
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Sandra Lavorel | Izaskun Casado-Arzuaga | Leah Oliver | Greg Guannel | Ehsan Rahini | Carl Walters | Jonathan Halama |
Contact Address
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Laboratoire d’Ecologie Alpine, UMR 5553 CNRS Université Joseph Fourier, BP 53, 38041 Grenoble Cedex 9, France | Plant Biology and Ecology Department, University of the Basque Country UPV/EHU, Campus de Leioa, Barrio Sarriena s/n, 48940 Leioa, Bizkaia, Spain | National Health and Environmental Research Effects Laboratory | The Nature Conservancy, Coral Gables, FL. USA | Environmental Sciences Research Institute, Shahid Beheshti University, Tehran, Iran | Fisheries Centre, University of British Columbia, Vancouver, British Columbia, British Columbia, Canada, V6T 1Z4 | U.S. Environmental Protection Agency, Corvallis, OR |
Contact Email
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sandra.lavorel@ujf-grenoble.fr | izaskun.casado@ehu.es | leah.oliver@epa.gov | greg.guannel@gmail.com | ehsanrahimi666@gmail.com | c.walters@oceans.ubc.ca | Halama.Jonathan@epa.gov |
EM ID
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EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
Summary Description
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ABSTRACT: "Here, we propose a new approach for the analysis, mapping and understanding of multiple ES delivery in landscapes. Spatially explicit single ES models based on plant traits and abiotic characteristics are combined to identify ‘hot’ and ‘cold’ spots of multiple ES delivery, and the land use and biotic determinants of such distributions. We demonstrate the value of this trait-based approach as compared to a pure land-use approach for a pastoral landscape from the central French Alps, and highlight how it improves understanding of ecological constraints to, and opportunities for, the delivery of multiple services. Vegetative height and leaf traits such as leaf dry matter content were response traits strongly influenced by land use and abiotic environment, with follow-on effects on several ecosystem properties (e.g., green biomass production), and could therefore be used as functional markers of ES." AUTHOR'S DESCRIPTION: "Variation in green biomass production was modelled using…traits community-weighted mean (CWM) and functional divergence (FD) and abiotic variables (continuous variables; trait + abiotic) following Diaz et al. (2007). …The comparison between this model and the land-use alone model identifies the need for site-based information beyond a land use or land cover proxy, and the comparison with the land use + abiotic model assesses the value of additional ecological (trait) information…Green biomass production for each pixel was calculated and mapped using model estimates for…regression coefficients on abiotic variables and traits. For each pixel these calculations were applied to mapped estimates of abiotic variables and trait CWM and FD. This step is critically novel as compared to a direct application of the model by Diaz et al. (2007) in that we explicitly modelled the responses of trait community-weighted means and functional divergences to environment prior to evaluating their effects on ecosystem properties. Such an approach is the key to the explicit representation of functional variation across the landscape, as opposed to the use of unique trait values within each land use (see Albert et al. 2010)." | ABSTRACT "This paper presents a method to quantify cultural ecosystem services (ES) and their spatial distribution in the landscape based on ecological structure and social evaluation approaches. The method aims to provide quantified assessments of ES to support land use planning decisions. A GIS-based approach was used to estimate and map the provision of recreation and aesthetic services supplied by ecosystems in a peri-urban area located in the Basque Country, northern Spain. Data of two different public participation processes (frequency of visits to 25 different sites within the study area and aesthetic value of different landscape units) were used to validate the maps. Three maps were obtained as results: a map showing the provision of recreation services, an aesthetic value map and a map of the correspondences and differences between both services. The data obtained in the participation processes were found useful for the validation of the maps. A weak spatial correlation was found between aesthetic quality and recreation provision services, with an overlap of the highest values for both services only in 7.2 % of the area. A consultation with decision-makers indicated that the results were considered useful to identify areas that can be targeted for improvement of landscape and recreation management." | AUTHOR'S DESCRIPTION: "In this exploratory comparison, stony coral condition was related to watershed LULC and LDI values. We also compared the capacity of other potential human activity indicators to predict coral reef condition using multivariate analysis." (294) | AUTHOR'S DESCRIPTION: "Natural habitats have the ability to protect coastal communities against the impacts of waves and storms, yet it is unclear how different habitats complement each other to reduce those impacts. Here, we investigate the individual and combined coastal protection services supplied by live corals on reefs, seagrass meadows, and mangrove forests during both non-storm and storm conditions, and under present and future sea-level conditions. Using idealized profiles of fringing and barrier reefs, we quantify the services supplied by these habitats using various metrics of inundation and erosion. We find that, together, live corals, seagrasses, and mangroves supply more protection services than any individual habitat or any combination of two habitats. Specifically, we find that, while mangroves are the most effective at protecting the coast under non-storm and storm conditions, live corals and seagrasses also moderate the impact of waves and storms, thereby further reducing the vulnerability of coastal regions. Also, in addition to structural differences, the amount of service supplied by habitats in our analysis is highly dependent on the geomorphic setting, habitat location and forcing conditions: live corals in the fringing reef profile supply more protection services than seagrasses; seagrasses in the barrier reef profile supply more protection services than live corals; and seagrasses, in our simulations, can even compensate for the long-term degradation of the barrier reef. Results of this study demonstrate the importance of taking integrated and place-based approaches when quantifying and managing for the coastal protection services supplied by ecosystems." | Abstract: ". ..we used the ESTIMAP model to improve the results of the Lonsdorf model. For this, we included the effects of roads, railways, rivers, wetlands, lakes, altitude, climate, and ecosystem boundaries in the ESTIMAP modeling and compared the results with the Lonsdorf model. The results of the Lonsdorf model showed that the majority of Iran had a very low potential for providing pollination service and only three percent of the northern and western parts of Iran had high potential. However, the results of the ESTIMAP model showed that 16% of Iran had a high potential to provide pollination that covers most of the northern and southern parts of the country. The results of the ESTIMAP model for pollination mapping in Iran showed the Lonsdorf model of estimating pollination service can be improved through considering other relevant factors." | ABSTRACT: " EcoSim II uses results from the Ecopath procedure for trophic mass-balance analysis to define biomass dynamics models for predicting temporal change in exploited ecosystems. Key populations can be repre- sented in further detail by using delay-difference models to account for both biomass and numbers dynamics. A major problem revealed by linking the population and biomass dynamics models is in representation of population responses to changes in food supply; simple proportional growth and reproductive responses lead to unrealistic predic- tions of changes in mean body size with changes in fishing mortality. EcoSim II allows users to specify life history mechanisms to avoid such unrealistic predictions: animals may translate changes in feed- ing rate into changes in reproductive rather than growth rates, or they may translate changes in food availability into changes in foraging time that in turn affects predation risk. These options, along with model relationships for limits on prey availabil- ity caused by predation avoidance tactics, tend to cause strong compensatory responses in modeled populations. It is likely that such compensatory responses are responsible for our inability to find obvious correlations between interacting trophic components in fisheries time-series data. But Eco- sim II does not just predict strong compensatory responses: it also suggests that large piscivores may be vulnerable to delayed recruitment collapses caused by increases in prey species that are in turn competitors/predators of juvenile piscivores " | ABSTRACT: "Coho salmon (Oncorhynchus kisutch) are highly sensitive to 6PPD-Quinone (6PPD-Q). Details of the hydrological and biogeochemical processes controlling spatial and temporal dynamics of 6PPD-Q fate and transport from points of deposition to receiving waters (e.g., streams, estuaries) are poorly understood. To understand the fate and transport of 6PPD and mechanisms leading to salmon mortality Visualizing Ecosystem Land Management Assessments (VELMA), an ecohydrological model developed by US Environmental Protection Agency (EPA), was enhanced to better understand and inform stormwater management planning by municipal, state, and federal partners seeking to reduce stormwater contaminant loads in urban streams draining to the Puget Sound National Estuary. This work focuses on the 5.5 km2 Longfellow Creek upper watershed (Seattle, Washington, United States), which has long exhibited high rates of acute urban runoff mortality syndrome in coho salmon. We present VELMA model results to elucidate these processes for the Longfellow Creek watershed across multiple scales–from 5-m grid cells to the entire watershed. Our results highlight hydrological and biogeochemical controls on 6PPD-Q flow paths, and hotspots within the watershed and its stormwater infrastructure, that ultimately impact contaminant transport to Longfellow Creek and Puget Sound. Simulated daily average 6PPD-Q and available observed 6PPD-Q peak in-stream grab sample concentrations (ng/L) corresponds within plus or minus 10 ng/L. Most importantly, VELMA’s high-resolution spatial and temporal analysis of 6PPD-Q hotspots provides a tool for prioritizing the locations, amounts, and types of green infrastructure that can most effectively reduce 6PPD-Q stream concentrations to levels protective of coho salmon and other aquatic species. " |
Specific Policy or Decision Context Cited
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None identified | Land management, ecosystem management, response to EU 2020 Biodiversity Strategy | Not applicable | Future rock lobster fisheries management | None reported | None | Not reported |
Biophysical Context
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Elevation ranges from 1552 to 2442 m, on predominately south-facing slopes | Northern Spain; Bizkaia region | nearshore; <1.5 km offshore; <12 m depth | barrier reef and fringing reef in nearshore coastal marine system | None additional | None, Ocean ecosystems | 6PPD deposition from vehicle tire wear particles. |
EM Scenario Drivers
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No scenarios presented | No scenarios presented | Not applicable | Reef type, Sea level increase, storm conditions, seagrass conditions, coral conditions, vegetation types and conditions | N/A | N/A | N/A |
EM ID
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EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
Method Only, Application of Method or Model Run
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Method + Application | Method + Application | Method + Application | Method + Application (multiple runs exist) View EM Runs | Method + Application | Method Only | Method + Application |
New or Pre-existing EM?
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New or revised model | New or revised model | New or revised model | New or revised model | Application of existing model | New or revised model | Application of existing model |
Related EMs (for example, other versions or derivations of this EM) described in ESML
EM ID
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EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
Document ID for related EM
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Doc-260 | None | None | None | Doc-432 | None | Doc-366 | Doc-423 | Doc-430 |
EM ID for related EM
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EM-66 | EM-68 | EM-69 | EM-70 | EM-71 | EM-79 | EM-80 | EM-81 | EM-82 | EM-83 | None | None | None | EM-939 | None | None |
EM Modeling Approach
EM ID
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EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
EM Temporal Extent
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2007-2009 | 2000 - 2007 | 2006-2007 | 2005-2013 | 2020 | Not applicable | 9/2020-6/2021 |
EM Time Dependence
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time-stationary | time-stationary | time-stationary | time-dependent | time-stationary | time-dependent | time-dependent |
EM Time Reference (Future/Past)
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Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | both | past time |
EM Time Continuity
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Not applicable | Not applicable | Not applicable | discrete | Not applicable |
discrete ?Comment:Modeller dependent |
discrete |
EM Temporal Grain Size Value
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Not applicable | Not applicable | Not applicable | 1 | Not applicable | 1 | 1 |
EM Temporal Grain Size Unit
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Not applicable | Not applicable | Not applicable | Second | Not applicable | Day | Day |
EM ID
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EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
Bounding Type
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Physiographic or Ecological | Geopolitical | Physiographic or Ecological | Geopolitical | Geopolitical | Other | Watershed/Catchment/HUC |
Spatial Extent Name
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Central French Alps | Bilbao Metropolitan Greenbelt | St.Croix, U.S. Virgin Islands | Coast of Belize | Iran | Not applicable | Longfellow creek |
Spatial Extent Area (Magnitude)
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10-100 km^2 | 100-1000 km^2 | 10-100 km^2 | 100-1000 km^2 | >1,000,000 km^2 | Not applicable | 1-10 km^2 |
EM ID
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EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
EM Spatial Distribution
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spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially lumped (in all cases) | spatially distributed (in at least some cases) |
spatially distributed (in at least some cases) ?Comment:Varies by inputs, but results are for areas of country |
spatially lumped (in all cases) | spatially lumped (in all cases) |
Spatial Grain Type
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area, for pixel or radial feature | area, for pixel or radial feature | Not applicable | length, for linear feature (e.g., stream mile) | area, for pixel or radial feature | Not applicable | Not applicable |
Spatial Grain Size
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20 m x 20 m | 2 m x 2 m | Not applicable | 1 meter | ha^2 | Not applicable | Not applicable |
EM ID
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EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
EM Computational Approach
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Analytic | Analytic | Analytic | Analytic | Numeric | Analytic | Analytic |
EM Determinism
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deterministic | deterministic | deterministic | deterministic | deterministic | deterministic | deterministic |
Statistical Estimation of EM
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EM ID
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EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
Model Calibration Reported?
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No | No | Yes | No | No | No | Yes |
Model Goodness of Fit Reported?
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Yes | No | Yes | No | No | No | No |
Goodness of Fit (metric| value | unit)
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None |
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None | None | None | None |
Model Operational Validation Reported?
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Yes | Yes | No |
No ?Comment:Used the SWAN model (see below for referenece) with Generation 1 or 2 wind-wave formulations to validate the wave development portion of the model. Booij N, Ris RC, Holthuijsen LH. A third-generation wave model for coastal regions 1. Model description and validation. J Geophys Res. American Geophysical Union; 1999;104: 7649?7666. |
No | Not applicable | Yes |
Model Uncertainty Analysis Reported?
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No | No | Yes | No | No | Not applicable | Unclear |
Model Sensitivity Analysis Reported?
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No | No | No | No | No | Not applicable | Unclear |
Model Sensitivity Analysis Include Interactions?
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Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable |
EM Locations, Environments, Ecology
Terrestrial location (Classification hierarchy: Continent > Country > U.S. State [United States only])
EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
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None |
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Comment:Model for Iran - no form preset id for country |
None |
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Marine location (Classification hierarchy: Realm > Region > Province > Ecoregion)
EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
None | None |
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None | None | None |
Centroid Lat/Long (Decimal Degree)
EM ID
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EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
Centroid Latitude
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45.05 | 43.25 | 17.75 | 18.63 | 32.29 | Not applicable | 47.55 |
Centroid Longitude
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6.4 | -2.92 | -64.75 | -88.22 | 53.68 | Not applicable | 122.37 |
Centroid Datum
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WGS84 | WGS84 | NAD83 | WGS84 | WGS84 | Not applicable | None provided |
Centroid Coordinates Status
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Provided | Provided | Estimated | Estimated | Estimated | Not applicable | Provided |
EM ID
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EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
EM Environmental Sub-Class
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Agroecosystems | Grasslands | Aquatic Environment (sub-classes not fully specified) | Rivers and Streams | Near Coastal Marine and Estuarine | Terrestrial Environment (sub-classes not fully specified) | Forests | Agroecosystems | Created Greenspace | Grasslands | Scrubland/Shrubland | Near Coastal Marine and Estuarine | Near Coastal Marine and Estuarine | Terrestrial Environment (sub-classes not fully specified) | Open Ocean and Seas | Rivers and Streams |
Specific Environment Type
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Subalpine terraces, grasslands, and meadows | none | stony coral reef | coral reefs | terrestrial land types | Pelagic | small stream |
EM Ecological Scale
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Not applicable | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class |
Scale of differentiation of organisms modeled
EM ID
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EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
EM Organismal Scale
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Community | Not applicable | Guild or Assemblage | Guild or Assemblage | Not applicable |
Other (Comment) ?Comment:Varied levels of taxonomic order |
Species |
Taxonomic level and name of organisms or groups identified
EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
None Available | None Available |
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None Available |
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EnviroAtlas URL
EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
CICES v 4.3 - Common International Classification of Ecosystem Services (Section > Division > Group > Class)
EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
None |
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<a target="_blank" rel="noopener noreferrer" href="https://www.epa.gov/eco-research/national-ecosystem-services-classification-system-nescs-plus">National Ecosystem Services Classification System (NESCS) Plus</a>
(Environmental Subclass > Ecological End-Product (EEP) > EEP Subclass > EEP Modifier)
EM-65 | EM-193 | EM-260 |
EM-542 ![]() |
EM-941 | EM-964 | EM-993 |
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Comment:Model identifies toxicant concentrations relative to the known LC50 for coho juveniles which is 95ng/L (Spromber and Scholz, 2011; |