EcoService Models Library (ESML)
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Compare EMs
Which comparison is best for me?EM Variables by Variable Role
One quick way to compare ecological models (EMs) is by comparing their variables. Predictor variables show what kinds of influences a model is able to account for, and what kinds of data it requires. Response variables show what information a model is capable of estimating.
This first comparison shows the names (and units) of each EM’s variables, side-by-side, sorted by variable role. Variable roles in ESML are as follows:
- Predictor Variables
- Time- or Space-Varying Variables
- Constants and Parameters
- Intermediate (Computed) Variables
- Response Variables
- Computed Response Variables
- Measured Response Variables
EM Variables by Category
A second way to use variables to compare EMs is by focusing on the kind of information each variable represents. The top-level categories in the ESML Variable Classification Hierarchy are as follows:
- Policy Regarding Use or Management of Ecosystem Resources
- Land Surface (or Water Body Bed) Cover, Use or Substrate
- Human Demographic Data
- Human-Produced Stressor or Enhancer of Ecosystem Goods and Services Production
- Ecosystem Attributes and Potential Supply of Ecosystem Goods and Services
- Non-monetary Indicators of Human Demand, Use or Benefit of Ecosystem Goods and Services
- Monetary Values
Besides understanding model similarities, sorting the variables for each EM by these 7 categories makes it easier to see if the compared models can be linked using similar variables. For example, if one model estimates an ecosystem attribute (in Category 5), such as water clarity, as a response variable, and a second model uses a similar attribute (also in Category 5) as a predictor of recreational use, the two models can potentially be used in tandem. This comparison makes it easier to spot potential model linkages.
All EM Descriptors
This selection allows a more detailed comparison of EMs by model characteristics other than their variables. The 50-or-so EM descriptors for each model are presented, side-by-side, in the following categories:
- EM Identity and Description
- EM Modeling Approach
- EM Locations, Environments, Ecology
- EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
EM Descriptors by Modeling Concepts
This feature guides the user through the use of the following seven concepts for comparing and selecting EMs:
- Conceptual Model
- Modeling Objective
- Modeling Context
- Potential for Model Linkage
- Feasibility of Model Use
- Model Certainty
- Model Structural Information
Though presented separately, these concepts are interdependent, and information presented under one concept may have relevance to other concepts as well.
EM Identity and Description
EM ID
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EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
EM Short Name
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ACRU, South Africa | Area and hotspots of flow regulation, South Africa | Area and hotspots of soil retention, South Africa | InVEST habitat quality | Evoland v3.5 (unbounded growth), Eugene, OR, USA | InVEST water yield, Xitiaoxi River basin, China | SAV occurrence, St. Louis River, MN/WI, USA | SIRHI, St. Croix, USVI | Nitrogen fixation rates, Guánica Bay, Puerto Rico | Chinook salmon value (household), Yaquina Bay, OR | WTP for a beach day, Massachusetts, USA | Floral resources on landfill sites, United Kingdom | WESP: Riparian & stream habitat, ID, USA | InVEST Coastal Vulnerability, New York, USA |
EM Full Name
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ACRU (Agricultural Catchments Research Unit), South Africa | Area and hotspots of water flow regulation, South Africa | Area and hotspots of soil retention, South Africa | InVEST (Integrated Valuation of Environmental Services and Tradeoffs) Habitat Quality | Evoland v3.5 (without urban growth boundaries), Eugene, OR, USA | InVEST (Integrated Valuation of Ecosystem Services and Tradeoffs) water yield, Xitiaoxi River basin, China | Predicting submerged aquatic vegetation occurrence, St. Louis River Estuary, MN & WI, USA | SIRHI (SImplified Reef Health Index), St. Croix, USVI | Nitrogen fixation rates, Guánica Bay, Puerto Rico, USA | Economic value of Chinook salmon per household method, Yaquina Bay, OR | Willingness to pay (WTP) for a beach day, Barnstable, Massachusetts, USA | Floral resources on landfill sites, East Midlands, United Kingdom | WESP: Riparian and stream habitat focus projects, ID, USA | InVEST Coastal Vulnerability, Jamaica Bay, New York, USA |
EM Source or Collection
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None | None | None |
InVEST ?Comment:From the Natural Capital Project website |
Envision | InVEST | US EPA | US EPA | US EPA | US EPA | US EPA | None | None | InVEST |
EM Source Document ID
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271 | 271 | 271 | 278 |
47 ?Comment:Doc 183 is a secondary source for the Evoland model. |
307 | 330 | 335 |
338 ?Comment:WE received a draft copy prior to journal publication that was agency reviewed. |
324 | 386 | 389 |
393 ?Comment:Additional data came from electronic appendix provided by author Chris Murphy. |
410 ?Comment:Sharp R, Tallis H, Ricketts T, Guerry A, Wood S, Chaplin-Kramer R, et al. InVEST User?s Guide. User Guide. Stanford (CA): The Natural Capital Project, Stanford University, University of Minnesota, The Nature Conservancy, World Wildlife Fund; 2015. |
Document Author
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Egoh, B., Reyers, B., Rouget, M., Richardson, D.M., Le Maitre, D.C., and van Jaarsveld, A.S. | Egoh, B., Reyers, B., Rouget, M., Richardson, D.M., Le Maitre, D.C., and van Jaarsveld, A.S. | Egoh, B., Reyers, B., Rouget, M., Richardson, D.M., Le Maitre, D.C., and van Jaarsveld, A.S. | Natural Capital Project | Guzy, M. R., Smith, C. L. , Bolte, J. P., Hulse, D. W. and Gregory, S. V. | Zhang C., Li, W., Zhang, B., and Liu, M. | Ted R. Angradi, Mark S. Pearson, David W. Bolgrien, Brent J. Bellinger, Matthew A. Starry, Carol Reschke | Yee, S. H., Dittmar, J. A., and L. M. Oliver | Amelia Smith, Susan Harrell Yee, Marc Russell, Jill Awkerman and William S. Fisher | Stephen J. Jordan, Timothy O'Higgins and John A. Dittmar | Lyon, Sarina F., Nathaniel H. Merrill, Kate K. Mulvaney, and Marisa J. Mazzotta | Tarrant S., J. Ollerton, M. L Rahman, J. Tarrant, and D. McCollin | Murphy, C. and T. Weekley | Hopper T. and M. S. Meixler |
Document Year
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2008 | 2008 | 2008 | 2014 | 2008 | 2012 | 2013 | 2014 | 2017 | 2012 | 2018 | 2013 | 2012 | 2016 |
Document Title
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Mapping ecosystem services for planning and management | Mapping ecosystem services for planning and management | Mapping ecosystem services for planning and management | Habitat Quality model - InVEST ver. 3.0 | Policy research using agent-based modeling to assess future impacts of urban expansion into farmlands and forests | Water yield of Xitiaoxi River basin based on InVEST modeling | Predicting submerged aquatic vegetation cover and occurrence in a Lake Superior estuary | Comparison of methods for quantifying reef ecosystem services: A case study mapping services for St. Croix, USVI | Linking ecosystem services supply to stakeholder concerns on both land and sea: An example from Guanica Bay watershed, Puerto Rico | Ecosystem Services of Coastal Habitats and Fisheries: Multiscale Ecological and Economic Models in Support of Ecosystem-Based Management | Valuing coastal beaches and closures using benefit transfer: An application to Barnstable, Massachusetts | Grassland restoration on landfill sites in the East Midlands, United Kingdom: An evaluation of floral resources and pollinating insects | Measuring outcomes of wetland restoration, enhancement, and creation in Idaho-- Assessing potential functions, values, and condition in a watershed context. | Modeling coastal vulnerability through space and time |
Document Status
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Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published | Peer reviewed and published |
Comments on Status
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Published journal manuscript | Published journal manuscript | Published journal manuscript | Published on Natural Capital Project website | Published journal manuscript | Published journal manuscript | Published journal manuscript | Published journal manuscript | Published journal manuscript | Published journal manuscript | Published journal manuscript | Published journal manuscript | Published report | Published journal manuscript |
EM ID
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EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
Not applicable | Not applicable | Not applicable | https://www.naturalcapitalproject.org/invest/ | http://evoland.bioe.orst.edu/ | https://www.naturalcapitalproject.org/invest/ | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | https://naturalcapitalproject.stanford.edu/software/invest-models/coastal-vulnerability | |
Contact Name
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Roland E Schulze | Benis Egoh | Benis Egoh | The Natural Capital Project | Michael R. Guzy | Li Wenhua | Ted R. Angradi | Susan H. Yee | Susan H. Yee | Stephen Jordan | Kate K, Mulvaney | Sam Tarrant | Chris Murphy | Thomas Hopper |
Contact Address
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School of Bioresources Engineering and Environmental Hydrology, University of Natal, South Africa | Water Resources Unit, Institute for Environment and Sustainability, European Commission - Joint Research Centre, Ispra, Italy | Water Resources Unit, Institute for Environment and Sustainability, European Commission - Joint Research Centre, Ispra, Italy | 371 Serra Mall Stanford University Stanford, CA 94305-5020 USA | Oregon State University, Dept. of Biological and Ecological Engineering | Institute of Geographic Sciences and Natural Resources Research, CAS, Beijing 100101, China | U.S. Environmental Protection Agency, Office of Research and Development, National Health and Environmental Effects Research Laboratory, Mid-Continent Ecology Division, 6201 Congdon Blvd., Duluth, MN 55804, USA | US EPA, Office of Research and Development, NHEERL, Gulf Ecology Division, Gulf Breeze, FL 32561, USA | U.S. Environmental Protection Agency, Gulf Ecology Division, Gulf Breeze, FL 32561, USA | U.S. EPA, Gulf Ecology Div., 1 Sabine Island Dr., Gulf Breeze, FL 32561, USA | Not reported | RSPB UK Headquarters, The Lodge, Sandy, Bedfordshire SG19 2DL, U.K. | Idaho Dept. Fish and Game, Wildlife Bureau, Habitat Section, Boise, ID | Not reported |
Contact Email
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schulzeR@nu.ac.za | Not reported | Not reported | invest@naturalcapitalproject.org | Not reported | liwh@igsnrr.ac.cn | angradi.theodore@epa.gov | yee.susan@epa.gov | yee.susan@epa.gov | jordan.steve@epa.gov | Mulvaney.Kate@EPA.gov | sam.tarrant@rspb.org.uk | chris.murphy@idfg.idaho.gov | Tjhop1123@gmail.com |
EM ID
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EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
Summary Description
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AUTHOR'S DESCRIPTION (Doc ID 272): "ACRU is a daily timestep, physical conceptual and multipurpose model structured to simulate impacts of land cover/ use change. The model can output, inter alia, components of runoff, irrigation supply and demand, reservoir water budgets as well as sediment and crop yields." AUTHOR'S DESCRIPTION (Doc ID 271): "We define the range of ecosystem services as areas of meaningful supply, similar to a species’ range or area of occupancy. The term ‘‘hotspots’’ was proposed by Norman Myers in the 1980s and refers to areas of high species richness, endemism and/or threat and has been widely used to prioritise areas for biodiversity conservation. Similarly, this study suggests that hotspots for ecosystem services are areas of critical management importance for the service. Here the term ecosystem service hotspot is used to refer to areas which provide large proportions of a particular service, and do not include measures of threat or endemism…The total benefit to people of water supply is a function of both the quantity and quality with the ecosystem playing a key role in the latter. However, due to the lack of suitable national scale data on water quality for quantifying the service, runoff was used as an estimate of the benefit where runoff is the total water yield from a watershed including surface and subsurface flow. This assumes that runoff is positively correlated with quality, which is the case in South Africa (Allanson et al., 1990)…In South Africa, water resources are mapped in water management areas called catchments (vs. watersheds) where a catchment is defined as the area of land that is drained by a single river system, including its tributaries (DWAF, 2004). There are 1946 quaternary (4th order) catchments in South Africa, the smallest is 4800 ha and the average size is 65,000 ha. Schulze (1997) modelled annual runoff for each quaternary catchment. During modelling of runoff, he used rainfall data collected over a period of more than 30 years, as well as data on other climatic factors, soil characteristics and grassland as the land cover. In this study, median annual simulated runoff was used as a measure of surface water supply. The volume of runoff per quaternary catchment was calculated for surface water supply. The range (areas with runoff of 30 million m^3 or more) and hotspots (areas with runoff of 70 million m^3 or more) were defined using a combination of statistics and expert inputs due to a lack of published thresholds in the literature." | AUTHOR'S DESCRIPTION: "We define the range of ecosystem services as areas of meaningful supply, similar to a species’ range or area of occupancy. The term ‘‘hotspots’’ was proposed by Norman Myers in the 1980s and refers to areas of high species richness, endemism and/or threat and has been widely used to prioritise areas for biodiversity conservation. Similarly, this study suggests that hotspots for ecosystem services are areas of critical management importance for the service. Here the term ecosystem service hotspot is used to refer to areas which provide large proportions of a particular service, and do not include measures of threat or endemism…Water flow regulation is a function of the storage and retention components of the water supply service (de Groot et al., 2002). The ability of a catchment to regulate flows is directly related to the volume of water that is retained or stored in the soil and underlying aquifers as moisture or groundwater; and the infiltration rate of water which replenishes the stored water (Kittredge, 1948; Farvolden, 1963). Groundwater contribution to surface runoff is the most direct measure of the water regulation function of a catchment. Data on the percentage contribution of groundwater to baseflows were obtained from DWAF (2005) per quaternary catchment and expressed as a percentage of total surface runoff, the range and hotspot being defined as areas with at least 10% and 30%, respectively (Colvin et al., 2007)." | AUTHOR'S DESCRIPTION: "We define the range of ecosystem services as areas of meaningful supply, similar to a species’ range or area of occupancy. The term ‘‘hotspots’’ was proposed by Norman Myers in the 1980s and refers to areas of high species richness, endemism and/or threat and has been widely used to prioritise areas for biodiversity conservation. Similarly, this study suggests that hotspots for ecosystem services are areas of critical management importance for the service. Here the term ecosystem service hotspot is used to refer to areas which provide large proportions of a particular service, and do not include measures of threat or endemism…Soil retention was modelled as a function of vegetation or litter cover and soil erosion potential. Schoeman et al. (2002) modelled soil erosion potential and derived eight erosion classes, ranging from low to severe erosion potential for South Africa. The vegetation cover was mapped by ranking vegetation types using expert knowledge of their ability to curb erosion. We used Schulze (2004) index of litter cover which estimates the soil surface covered by litter based on observations in a range of grasslands, woodlands and natural forests. According to Quinton et al. (1997) and Fowler and Rockstrom (2001) soil erosion is slightly reduced with about 30%, significantly reduced with about 70% vegetation cover. The range of soil retention was mapped by selecting all areas that had vegetation or litter cover of more than 30% for both the expert classified vegetation types and litter accumulation index within areas with moderate to severe erosion potential. The hotspot was mapped as areas with severe erosion potential and vegetation/litter cover of at least 70% where maintaining the cover is essential to prevent erosion. An assumption was made that the potential for this service is relatively low in areas with little natural vegetation or litter cover." | Please note: This ESML entry describes an InVEST model version that was current as of 2014. More recent versions may be available at the InVEST website. AUTHORS DESCRIPTION: "The InVEST habitat quality model combines information on LULC and threats to biodiversity to produce habitat quality maps. This approach generates two key sets of information that are useful in making an initial assessment of conservation needs: the relative extent and degradation of different types of habitat types in a region and changes across time. This approach further allows rapid assessment of the status of and change in a proxy for more detailed measures of biodiversity status. If habitat changes are taken as representative of genetic, species, or ecosystem changes, the user is assuming that areas with high quality habitat will better support all levels of biodiversity and that decreases in habitat extent and quality over time means a decline in biodiversity persistence, resilience, breadth and depth in the area of decline. The habitat rarity model indicates the extent and pattern of natural land cover types on the current or a potential future landscape vis-a-vis the extent of the same natural land cover types in some baseline period. Rarity maps allow users to create a map of the rarest habitats on the landscape relative to the baseline chosen by the user to represent the mix of habitats on the landscape that is most appropriate for the study area’s native biodiversity. The model requires basic data that are available virtually everywhere in the world, making it useful in areas for which species distribution data are poor or lacking altogether. Extensive occurrence (presence/absence) data may be available in many places for current conditions. However, modeling the change in occurrence, persistence, or vulnerability of multiple species under future conditions is often impossible or infeasible. While a habitat approach leaves out the detailed species occurrence data available for current conditions, several of its components represent advances in functionality over many existing biodiversity conservation planning tools. The most significant is the ability to characterize the sensitivity of habitats types to various threats. Not all habitats are affected by all threats in the same way, and the InVEST model accounts for this variability. Further, the model allows users to estimate the relative impact of one threat over another so that threats that are more damaging to biodiversity persistence on the landscape can be represented as such. For example, grassland could be particularly sensitive to threats generated by urban areas yet moderately sensitive to threats generated by roads. In addition, the distance over which a threat will degrade natural systems can be incorporated into the model. Model assessment of the current landscape can be used as an input to a coarse-filter assessment of current conservation needs and opportunities. Model assessment of pote | **Note: A more recent version of this model exists. See Related EMs below for links to related models/applications.** ABSTRACT: "Spatially explicit agent-based models can represent the changes in resilience and ecological services that result from different land-use policies…This type of analysis generates ensembles of alternate plausible representations of future system conditions. User expertise steers interactive, stepwise system exploration toward inductive reasoning about potential changes to the system. In this study, we develop understanding of the potential alternative futures for a social-ecological system by way of successive simulations that test variations in the types and numbers of policies. The model addresses the agricultural-urban interface and the preservation of ecosystem services. The landscape analyzed is at the junction of the McKenzie and Willamette Rivers adjacent to the cities of Eugene and Springfield in Lane County, Oregon." AUTHOR'S DESCRIPTION: "Two general scenarios for urban expansion were created to set the bounds on what might be possible for the McKenzie-Willamette study area. One scenario, fish conservation, tried to accommodate urban expansion, but gave the most weight to policies that would produce resilience and ecosystem services to restore threatened fish populations. The other scenario, unconstrained development, reversed the weighting. The 35 policies in the fish conservation scenario are designed to maintain urban growth boundaries (UGB), accommodate human population growth through increased urban densities, promote land conservation through best-conservation practices on agricultural and forest lands, and make rural land-use conversions that benefit fish. In the unconstrained development scenario, 13 policies are mainly concerned with allowing urban expansion in locations desired by landowners. Urban expansion in this scenario was not constrained by the extent of the UGB, and the policies are not intended to create conservation land uses." | Please note: This ESML entry describes a specific, published application of an InVEST model. Different versions (e.g. different tiers) or more recent versions of this model may be available at the InVEST website. ABSTRACT: "A water yield model based on InVEST was employed to estimate water runoff in the Xitiaoxi River basin…In order to test model accuracy the natural runoff of Xitiaoxi River was estimated based on linear regression relation of rainfall-runoff in a 'reference period'." AUTHOR'S DESCRIPTION: "The water yield model is based on the Budyko curve (1974) and annual precipitation…Water yield models require land use and land cover, precipitation, average annual potential evapotranspiration, soil depth, plant available water content, watersheds and sub-watersheds as well as a biophysical table reflecting the attributes of each land use and land cover." | ABSTRACT: “Submerged aquatic vegetation (SAV) provides the biophysical basis for multiple ecosystem services in Great Lakes estuaries. Understanding sources of variation in SAV is necessary for sustainable management of SAV habitat. From data collected using hydroacoustic survey methods, we created predictive models for SAV in the St. Louis River Estuary (SLRE) of western Lake Superior. The dominant SAV species in most areas of the estuary was American wild celery (Vallisneria americana Michx.)…” AUTHOR’S DESCRIPTION: “The SLRE is a Great Lakes “rivermouth” ecosystem as defined by Larson et al. (2013). The 5000-ha estuary forms a section of the state border between Duluth, Minnesota and Superior, Wisconsin…In the SLRE, SAV beds are often patchy, turbidity varies considerably among areas (DeVore, 1978) and over time, and the growing season is short. Given these conditions, hydroacoustic survey methods were the best option for generating the extensive, high resolution data needed for modeling. From late July through mid September in 2011, we surveyed SAV in Allouez Bay, part of Superior Bay, eastern half of St. Louis Bay, and Spirit Lake…We used the measured SAV percent cover at the location immediately previous to each useable record location along each transect as a lag variable to correct for possible serial autocorrelation of model error. SAV percent cover, substrate parameters, corrected depth, and exposure and bed slope data were combined in Arc-GIS...We created logistic regression models for each area of the SLRE to predict the probability of SAV being present at each report location. We created models for the training data set using the Logistic procedure in SAS v.9.1 with step wise elimination (?=0.05). Plots of cover by depth for selected predictor values (Supplementary Information Appendix C) suggested that interactions between depth and other predictors were likely to be significant, and so were included in regression models. We retained the main effect if their interaction terms were significant in the model. We examined the performance of the models using the area under the receiver operating characteristic (AUROC) curve. AUROC is the probability of concordance between random pairs of observations and ranges from 0.5 to 1 (Gönen, 2006). We cross-validated logistic occurrence models for their ability to classify correctly locations in the validation (holdout) dataset and in the Superior Bay dataset… Model performance, as indicated by the area under the receiver operating characteristic (AUROC) curve was >0.8 (Table 3). Assessed accuracy of models (the percent of records where the predicted probability of occurrence and actual SAV presence or absence agreed) for split datasets was 79% for Allouez Bay, 86% for St. Louis Bay, and 78% for Spirit Lake." | ABSTRACT: "...We investigated and compared a number of existing methods for quantifying ecological integrity, shoreline protection, recreational opportunities, fisheries production, and the potential for natural products discovery from reefs. Methods were applied to mapping potential ecosystem services production around St. Croix, U.S. Virgin Islands. Overall, we found that a number of different methods produced similar predictions." AUTHOR'S DESCRIPTION: "A number of methods have been developed for linking biophysical attributes of reef condition, such as reef structural complexity, fish biomass, or species richness, to provisioning of ecosystem goods and services (Principe et al., 2012). We investigated the feasibility of using existing methods and data for mapping production of reef ecosystem goods and services. We applied these methods toward mapping potential ecosystem goods and services production in St. Croix, U.S. Virgin Islands (USVI)...For each of the five categories of ecosystem services, we chose a suite of models and indices for estimating potential production based on relative ease of implementation, consisting of well-defined parameters, and likely availability of input data, to maximize potential for transferability to other locations. For each method, we assembled the necessary reef condition and environmental data as spatial data layers for St. Croix (Table1). The coastal zone surrounding St. Croix was divided into 10x10 m grid cells, and production functions were applied to quantify ecosystem services provisioning in each grid cell...A number of indicators have been proposed for measuring reef integrity, defined as the capacity to maintain healthy function and retention of diversity (Turner et al., 2000). The Simplified Integrated Reef Health Index (SIRHI) combines four attributes of reef condition into a single index: SIRHI = ΣiGi where Gi are the grades on a scale of 1 to 5 for four key reef attributes: percent coral cover, percent macroalgal cover, herbivorous fish biomass, and commercial fish biomass (Table2; Healthy Reefs Initiative, 2010). For a number of coral reef condition attributes, including fish richness, coral richness, and reef structural complexity, available data were point surveys from field monitoring by the US Environmental Protection Agency (see Oliver et al. (2011)) or the NOAA Caribbean Coral Reef Ecosystem Monitoring Program (see Pittman et al. (2008)). To generate continuous maps of coral condition for St. Croix, we fitted regression tree models to point survey data for St. Croix and then used models to predict reef condition in non-sampled locations (Fig. 1). In general, we followed the methods of Pittman et al. (2007) which generated predictive models for fish richness using readily available benthic habitat maps and bathymetry data. Because these models rely on readily available data (benthic habitat maps and bathymetry data), the models have the potential for high transferability to other locati | AUTHOR'S DESCRIPTION: " …In Guánica Bay watershed, Puerto Rico, deforestation and drainage of a large lagoon have led to sediment, contaminant, and nutrient transport into the bay, resulting in declining quality of coral reefs. A watershed management plan is currently being implemented to restore reefs through a variety of proposed actions…After the workshops, fifteen indicators of terrestrial ecosystem services in the watershed and four indicators in the coastal zone were identified to reflect the wide range of stakeholder concerns that could be impacted by management decisions. Ecosystem service production functions were applied to quantify and map ecosystem services supply in the Guánica Bay watershed, as well as an additional highly engineered upper multi-watershed area connected to the lower watershed via a series of reservoirs and tunnels,…” AUTHOR''S DESCRIPTION: "The U.S. Coral Reef Task Force (CRTF), a collaboration of federal, state and territorial agencies, initiated a program in 2009 to better incorporate land-based sources of pollution and socio-economic considerations into watershed strategies for coral reef protection (Bradley et al., 2016)...Baseline measures for relevant ecosystem services were calculated by parameterizing existing methods, largely based on land cover (Egoh et al., 2012; Martinez- Harms and Balvanera, 2012), with relevant rates of ecosystem services production for Puerto Rico, and applying them to map ecosystem services supply for the Guánica Bay Watershed...The Guánica Bay watershed is a highly engineered watershed in southwestern Puerto Rico, with a series of five reservoirs and extensive tunnel systems artificially connecting multiple mountainous sub-watersheds to the lower watershed of the Rio Loco, which itself is altered by an irrigation canal and return drainage ditch that diverts water through the Lajas Valley (PRWRA, 1948)...For each objective, a translator of ecosystem services production, i.e., ecological production function, was used to quantify baseline measurements of ecosystem services supply from land use/land cover (LULC) maps for watersheds across Puerto Rico...Two additional metrics, nitrogen fixation and rates of carbon sequestration into soil and sediment, were also calculated as potential measures of soil quality and agricultural productivity. Carbon sequestration and nitrogen fixation rates were assigned to each land cover class" | ABSTRACT:"Critical habitats for fish and wildlife are often small patches in landscapes, e.g., aquatic vegetation beds, reefs, isolated ponds and wetlands, remnant old-growth forests, etc., yet the same animal populations that depend on these patches for reproduction or survival can be extensive, ranging over large regions, even continents or major ocean basins. Whereas the ecological production functions that support these populations can be measured only at fine geographic scales and over brief periods of time, the ecosystem services (benefits that ecosystems convey to humans by supporting food production, water and air purification, recreational, esthetic, and cultural amenities, etc.) are delivered over extensive scales of space and time. These scale mismatches are particularly important for quantifying the economic values of ecosystem services. Examples can be seen in fish, shellfish, game, and bird populations. Moreover, there can be wide-scale mismatches in management regimes, e.g., coastal fisheries management versus habitat management in the coastal zone. We present concepts and case studies linking the production functions (contributions to recruitment) of critical habitats to commercial and recreational fishery values by combining site specific research data with spatial analysis and population models. We present examples illustrating various spatial scales of analysis, with indicators of economic value, for recreational Chinook (Oncorhynchus tshawytscha) salmon fisheries in the U.S. Pacific Northwest (Washington and Oregon) and commercial blue crab (Callinectes sapidus) and penaeid shrimp fisheries in the Gulf of Mexico. | ABSTRACT: "Each year, millions of Americans visit beaches for recreation, resulting in significant social welfare benefits and economic activity. Considering the high use of coastal beaches for recreation, closures due to bacterial contamination have the potential to greatly impact coastal visitors and communities. We used readily-available information to develop two transferable models that, together, provide estimates for the value of a beach day as well as the lost value due to a beach closure. We modeled visitation for beaches in Barnstable, Massachusetts on Cape Cod through panel regressions to predict visitation by type of day, for the season, and for lost visits when a closure was posted. We used a meta-analysis of existing studies conducted throughout the United States to estimate a consumer surplus value of a beach visit of around $22 for our study area, accounting for water quality at beaches by using past closure history. We applied this value through a benefit transfer to estimate the value of a beach day, and combined it with lost town revenue from parking to estimate losses in the event of a closure. The results indicate a high value for beaches as a public resource and show significant losses to the town when beaches are closed due to an exceedance in bacterial concentrations." AUTHOR'S DESCRIPTION: "We used existing studies in a meta-analysis to estimate appropriate benefit transfer values of consumer surplus per beach visit for Barnstable. The studies we include in the model are for beaches across the United States, allowing the metaregression model to be more broadly applicable to other beaches and for values to be adjusted based on appropriate site attributes...To identify relevant studies, we selected 25 studies of beach use and swimming from the Recreation Use Values Database (RUVD), where consumer surplus values are presented as value per day in 2016 dollars...We added beach length and history of closures to contextualize the model for our application by proxying water quality and site quality." Equation 1, page 11, provides the meta-regression. | ABSTRACT: "...Restored landfill sites are a significant potential reserve of semi-natural habitat, so their conservation value for supporting populations of pollinating insects was here examined by assessing whether the plant and pollinator assemblages of restored landfill sites are comparable to reference sites of existing wildlife value. Floral characteristics of the vegetation and the species richness and abundance of flower-visiting insect assemblages were compared between nine pairs of restored landfill sites and reference sites in the East Midlands of the United Kingdom, using standardized methods over two field seasons. …" AUTHOR'S DESCRIPTION: "The selection criteria for the landfill sites were greater than or equal to 50% of the site restored (to avoid undue influence from ongoing landfilling operations), greater than or equal to 0.5 ha in area and restored for greater than or equal to 4 years to allow establishment of vegetation. Comparison reference sites were the closest grassland sites of recognized nature conservation value, being designated as either Local Nature Reserves (LNRs) or Sites of Special Scientific Interest (SSSI)…All sites were surveyed three times each during the fieldwork season, in Spring, Summer, and Autumn. Paired sites were sampled on consecutive days whenever weather conditions permitted to reduce temporal bias. Standardized plant surveys were used (Dicks et al. 2002; Potts et al. 2006). Transects (100 × 2m) were centered from the approximate middle of the site and orientated using randomized bearing tables. All flowering plants were identified to species level… A “floral cover” method to represent available floral resources was used which combines floral abundance with inflorescence size. Mean area of the floral unit from above was measured for each flowering plant species and then multiplied by their frequencies." "Insect pollinated flowering plant species composition and floral abundance between sites by type were represented by non-metric multidimensional scaling (NMDS)...This method is sensitive to showing outliers and the distance between points shows the relative similarity (McCune & Grace 2002; Ollerton et al. 2009)." (This data is not entered into ESML) | A wetland restoration monitoring and assessment program framework was developed for Idaho. The project goal was to assess outcomes of substantial governmental and private investment in wetland restoration, enhancement and creation. The functions, values, condition, and vegetation at restored, enhanced, and created wetlands on private and state lands across Idaho were retrospectively evaluated. Assessment was conducted at multiple spatial scales and intensities. Potential functions and values (ecosystem services) were rapidly assessed using the Oregon Rapid Wetland Assessment Protocol. Vegetation samples were analyzed using Floristic Quality Assessment indices from Washington State. We compared vegetation of restored, enhanced, and created wetlands with reference wetlands that occurred in similar hydrogeomorphic environments determined at the HUC 12 level. | ABSTRACT: "Coastal ecosystems experience a wide range of stressors including wave forces, storm surge, sea-level rise, and anthropogenic modification and are thus vulnerable to erosion. Urban coastal ecosystems are especially important due to the large populations these limited ecosystems serve. However, few studies have addressed the issue of urban coastal vulnerability at the landscape scale with spatial data that are finely resolved. The purpose of this study was to model and map coastal vulnerability and the role of natural habitats in reducing vulnerability in Jamaica Bay, New York, in terms of nine coastal vulnerability metrics (relief, wave exposure, geomorphology, natural habitats, exposure, exposure with no habitat, habitat role, erodible shoreline, and surge) under past (1609), current (2015), and future (2080) scenarios using InVEST 3.2.0. We analyzed vulnerability results both spatially and across all time periods, by stakeholder (ownership) and by distance to damage from Hurricane Sandy. We found significant differences in vulnerability metrics between past, current and future scenarios for all nine metrics except relief and wave exposure…" |
Specific Policy or Decision Context Cited
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None identified | None identified | None identified | None identified | Authors Description: " By policy, we mean land management options that span the domains of zoning, agricultural and forest production, environmental protection, and urban development, including the associated regulations, laws, and practices. The policies we used in our SES simulations include urban containment policies…We also used policies modeled on agricultural practices that affect ecoystem services and capital…" | None identified | None identified | None identified | None provided | None identified | Economic value of protecting coastal beach water quality from contamination caused closures. | None identified | None identified | None identified |
Biophysical Context
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Semi-arid environment. Rainfall varies geographically from less than 50 to about 3000 mm per year (annual mean 450 mm). Soils are mostly very shallow with limited irrigation potential. | Semi-arid environment. Rainfall varies geographically from less than 50 to about 3000 mm per year (annual mean 450 mm). Soils are mostly very shallow with limited irrigation potential. | Semi-arid environment. Rainfall varies geographically from less than 50 to about 3000 mm per year (annual mean 450 mm). Soils are mostly very shallow with limited irrigation potential. | Not applicable | No additional description provided | Mean elevation of 266 m, with southwestern mountainous area. Subtropical monsoon climate. Annual average temperature of 12.2-15.6 °C. Annual mean precipitation is 1500 mm, and over 70% of rainfall occurs in the flood season (Apr-Oct). | submerged aquatic vegetation | No additional description provided | No additional description provided | Yaquina Bay estuary | Four separate beaches within the community of Barnstable | No additional description provided | restored, enhanced and created wetlands | Jamaica Bay, New York, situated on the southern shore of Long Island, and characterized by extensive coastal ecosystems in the central bay juxtaposed with a largely urbanized shoreline containing fragmented and fringing coastal habitat. |
EM Scenario Drivers
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No scenarios presented | No scenarios presented | No scenarios presented |
Potential land Use Land Class (LULC) future and baseline ?Comment:model requires current landuse but can compare to baseline (prior to intensive management of the land) and potential future landuse. These are the two scenarios suggested in the documentation. |
Three scenarios without urban growth boundaries, and with various combinations of unconstrainted development, fish conservation, and agriculture and forest reserves. | No scenarios presented | No scenarios presented | No scenarios presented | No scenarios presented | No scenarios presented | No scenarios presented | No scenarios presented | Sites, function or habitat focus | Past (1609), current (2015), and future (2080) scenarios |
EM ID
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EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
Method Only, Application of Method or Model Run
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Method + Application | Method + Application | Method + Application | Method Only | Method + Application (multiple runs exist) View EM Runs | Method + Application | Method + Application | Method + Application | Method + Application | Method + Application | Method + Application | Method + Application (multiple runs exist) View EM Runs | Method + Application (multiple runs exist) View EM Runs | Method + Application (multiple runs exist) View EM Runs |
New or Pre-existing EM?
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Application of existing model | New or revised model | New or revised model | New or revised model | New or revised model | Application of existing model | New or revised model | Application of existing model | Application of existing model | New or revised model | New or revised model | New or revised model | Application of existing model | Application of existing model |
Related EMs (for example, other versions or derivations of this EM) described in ESML
EM Modeling Approach
EM ID
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EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
EM Temporal Extent
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1950-1993 | Not reported | Not reported | Not applicable | 1990-2050 | 2003-2007 | 2010 - 2012 | 2006-2007, 2010 | 1978 - 2009 | 2003-2008 | July 1, 2011 to June 31, 2016 | 2007-2008 | 2010-2011 | 1609-2080 |
EM Time Dependence
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time-dependent | time-stationary | time-stationary | time-stationary | time-dependent | time-stationary | time-stationary | time-stationary | time-stationary | time-stationary | time-stationary | time-stationary | time-dependent | time-stationary |
EM Time Reference (Future/Past)
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future time | Not applicable | Not applicable | Not applicable | future time | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | past time | Not applicable |
EM Time Continuity
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discrete | Not applicable | Not applicable | Not applicable | discrete | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable |
EM Temporal Grain Size Value
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1 | Not applicable | Not applicable | Not applicable | 2 | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable |
EM Temporal Grain Size Unit
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Day | Not applicable | Not applicable | Not applicable | Year | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable |
EM ID
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EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
Bounding Type
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Geopolitical | Geopolitical | Geopolitical | No location (no locational reference given) | Geopolitical | Watershed/Catchment/HUC | Physiographic or ecological | Physiographic or ecological | Watershed/Catchment/HUC | Geopolitical | Physiographic or ecological | Multiple unrelated locations (e.g., meta-analysis) | Multiple unrelated locations (e.g., meta-analysis) | Physiographic or ecological |
Spatial Extent Name
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South Africa | South Africa | South Africa | Not applicable | Junction of McKenzie and Willamette Rivers, adjacent to the cities of Eugene and Springfield, Lane Co., Oregon, USA | Xitiaoxi River basin | St. Louis River Estuary | Coastal zone surrounding St. Croix | Guanica Bay watershed | Pacific Northwest | Barnstable beaches (Craigville Beach, Kalmus Beach, Keyes Memorial Beach, and Veteran’s Park Beach) | East Midlands | Wetlands in idaho | Jamaica Bay, Long Island, New York |
Spatial Extent Area (Magnitude)
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>1,000,000 km^2 | >1,000,000 km^2 | >1,000,000 km^2 | Not applicable | 10-100 km^2 | 1000-10,000 km^2. | 10-100 km^2 | 100-1000 km^2 | 100-1000 km^2 | >1,000,000 km^2 | 10-100 ha | 1000-10,000 km^2. | 100,000-1,000,000 km^2 | 10-100 km^2 |
EM ID
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EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
EM Spatial Distribution
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spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially distributed (in at least some cases) |
spatially distributed (in at least some cases) ?Comment:BH: Each individual transect?s data was parceled into location reports, and that each report?s ?quadrat? area was dependent upon the angle of the hydroacoustic sampling beam. The spatial grain is 0.07 m^2, 0.20 m^2 and 0.70 m^2 for depths of 1 meter, 2 meters and 3 meters, respectively. |
spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially lumped (in all cases) | spatially distributed (in at least some cases) | spatially distributed (in at least some cases) | spatially lumped (in all cases) |
spatially distributed (in at least some cases) ?Comment:by coastal segment |
Spatial Grain Type
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other (specify), for irregular (e.g., stream reach, lake basin) | other (specify), for irregular (e.g., stream reach, lake basin) | other (specify), for irregular (e.g., stream reach, lake basin) | area, for pixel or radial feature | area, for pixel or radial feature | area, for pixel or radial feature | area, for pixel or radial feature | area, for pixel or radial feature | other (specify), for irregular (e.g., stream reach, lake basin) | Not applicable | length, for linear feature (e.g., stream mile) | other (specify), for irregular (e.g., stream reach, lake basin) | Not applicable | length, for linear feature (e.g., stream mile) |
Spatial Grain Size
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Distributed by catchments with average size of 65,000 ha | Distributed by catchments with average size of 65,000 ha | Distributed across catchments with average size of 65,000 ha | LULC pixel size | varies | Not reported | 0.07 m^2 to 0.70 m^2 | 10 m x 10 m | HUC | Not applicable | by beach site | multiple unrelated locations | Not applicable | 80 m |
EM ID
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EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
EM Computational Approach
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Numeric | Analytic | Analytic | Analytic | Numeric | Analytic | Analytic | Analytic | Analytic | Analytic | Analytic | Analytic | Numeric | Analytic |
EM Determinism
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deterministic | deterministic | deterministic | deterministic | stochastic | deterministic | deterministic | deterministic | deterministic | deterministic | deterministic | deterministic | deterministic | deterministic |
Statistical Estimation of EM
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EM ID
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EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
Model Calibration Reported?
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No | No | No | Not applicable | Unclear | Yes | Yes | Yes | No | No | Yes | Not applicable | No | No |
Model Goodness of Fit Reported?
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No | No | No | Not applicable | No | No | Yes | No | No | No | Yes | Not applicable | No | No |
Goodness of Fit (metric| value | unit)
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None | None | None | None | None | None |
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None | None | None |
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None | None | None |
Model Operational Validation Reported?
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No | No | No | Not applicable | No | No | Yes | Yes | No | Yes | No | Not applicable | No | No |
Model Uncertainty Analysis Reported?
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No | No | No | Not applicable | No | No | No | No | No | No | No | Not applicable | No | No |
Model Sensitivity Analysis Reported?
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No | No | No | Not applicable | No | Yes | No | No | No | No |
Yes ?Comment:p-values of <0.05 and <0.01 provided for regression coefficient explanatory variables. |
Not applicable | No | No |
Model Sensitivity Analysis Include Interactions?
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Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | No | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable |
EM Locations, Environments, Ecology
Terrestrial location (Classification hierarchy: Continent > Country > U.S. State [United States only])
EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
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None |
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Marine location (Classification hierarchy: Realm > Region > Province > Ecoregion)
EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
None | None | None | None | None | None | None |
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None |
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None | None |
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Centroid Lat/Long (Decimal Degree)
EM ID
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EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
Centroid Latitude
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-30 | -30 | -30 | -9999 | 44.11 | 30.55 | 46.72 | 17.73 | 17.96 | 44.62 | 41.64 | 52.22 | 44.06 | 40.61 |
Centroid Longitude
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25 | 25 | 25 | -9999 | -123.09 | 119.5 | -96.13 | -64.77 | -67.02 | -124.02 | -70.29 | -0.91 | -114.69 | -73.84 |
Centroid Datum
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WGS84 | WGS84 | WGS84 | Not applicable | WGS84 | WGS84 | WGS84 | WGS84 | WGS84 | WGS84 | WGS84 | WGS84 | WGS84 | WGS84 |
Centroid Coordinates Status
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Estimated | Estimated | Estimated | Not applicable | Estimated | Provided | Estimated | Estimated | Estimated | Estimated | Estimated | Estimated | Estimated | Provided |
EM ID
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EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
EM Environmental Sub-Class
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Rivers and Streams | Ground Water | Terrestrial Environment (sub-classes not fully specified) | Rivers and Streams | Ground Water | Terrestrial Environment (sub-classes not fully specified) | Terrestrial Environment (sub-classes not fully specified) | Terrestrial Environment (sub-classes not fully specified) | Rivers and Streams | Forests | Agroecosystems | Created Greenspace | Rivers and Streams | Terrestrial Environment (sub-classes not fully specified) | Aquatic Environment (sub-classes not fully specified) | Rivers and Streams | Inland Wetlands | Lakes and Ponds | Near Coastal Marine and Estuarine | Inland Wetlands | Forests | Agroecosystems | Created Greenspace | Grasslands | Scrubland/Shrubland | Barren | Near Coastal Marine and Estuarine | Terrestrial Environment (sub-classes not fully specified) | Near Coastal Marine and Estuarine | Created Greenspace | Grasslands | Inland Wetlands | Near Coastal Marine and Estuarine |
Specific Environment Type
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Not reported | Not reported | Not reported | Not applicable | Agricultural-urban interface at river junction | Watershed | Freshwater estuarine system | Coral reefs | Tropical terrestrial | Yaquina Bay estuary and ocean | Saltwater beach | restored landfills and grasslands | created, restored and enhanced wetlands | Coastal |
EM Ecological Scale
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Ecological scale is coarser than that of the Environmental Sub-class | Ecological scale is coarser than that of the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class | Ecological scale is finer than that of the Environmental Sub-class | Ecological scale corresponds to the Environmental Sub-class |
Scale of differentiation of organisms modeled
EM ID
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EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
EM Organismal Scale
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Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Not applicable | Guild or Assemblage | Not applicable | Other (multiple scales) | Not applicable | Individual or population, within a species | Not applicable | Not applicable |
Taxonomic level and name of organisms or groups identified
EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
None Available | None Available | None Available | None Available |
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None Available | None Available |
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None Available |
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None Available |
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None Available | None Available |
EnviroAtlas URL
EM Ecosystem Goods and Services (EGS) potentially modeled, by classification system
CICES v 4.3 - Common International Classification of Ecosystem Services (Section > Division > Group > Class)
EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
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<a target="_blank" rel="noopener noreferrer" href="https://www.epa.gov/eco-research/national-ecosystem-services-classification-system-nescs-plus">National Ecosystem Services Classification System (NESCS) Plus</a>
(Environmental Subclass > Ecological End-Product (EEP) > EEP Subclass > EEP Modifier)
EM-84 | EM-85 | EM-86 | EM-143 |
EM-333 ![]() |
EM-344 | EM-414 | EM-418 | EM-432 | EM-604 | EM-682 |
EM-697 ![]() |
EM-718 ![]() |
EM-851 ![]() |
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None | None | None |
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None | None |
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None | None | None |